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Yacare Caiman (Caiman yacare) with butterfly (Paulogramma pyracmon) resting on its snout, Pantanal, Brazil
Bowhead Whale (Balaena mysticetus) rubbing off flaking skin on the ocean bottom, east coast Baffin island, Qikiqtarjuaq, Nunavut, Canada, May
Female Jaguar (Panthera onca) playing with her cub, captive, occurs in Southern and Central America
Black Lemur (Eulemur macaco) male, Nosy Komba, Madagascar
Pika (Ochotona princeps) in scree rock pile, Sheepeaters Cliff, Yellowstone National Park, Wyoming, USA, September
Grey squirrel (Sciurus carolinensis) adult and baby, against white background
Black-and-white kitten standing, against white background DIGITALLY ENHANCED
Dalmatian dog, Jack, 5 years, with one black ear, lying with head up, against white background
Harvest mouse (Micromys minutus) on wheat stem feeding, Devon, UK, July. Captive
American Red fox (Vulpes vulpes fulva) baby leaping on its disinterested mother. Grand Teton National Park, Wyoming, USA, May
Abandoned house full of sand. Kolmanskop Ghost Town, an old diamond-mining town where shifting sand dunes have encroached abandoned houses, Namib Desert Namibia, October 2013
Pine marten (Martes martes) juvenile male sitting on branch of a dead tree outside an old black woodpecker (Dryocopus martius) nest hole. Molde, Central Norway, September
Northern raccoon (Procyon lotor), group standing on branch, captive
African horned viper (Cerastes cerastes), captive
Paradise Riflebird (Ptiloris paradiseus) young males size each other up between practice displays. Australia
Woodcock orchid (Ophrys cornuta / scolopax) with mating Black veined white butterfliesWoodcock orchid (Ophrys cornuta/scolopax) with mating Black veined white butterflies (Aporia crataegi), Bela Reka, Eastern Rhodope Mountains, Bulgaria, May 2013
Lionesses (Panthera leo) grooming each other, Masai-Mara Game Reserve, Kenya
The narrow Nes Canyon, a fault filled with fresh water in the rift valley between the Eurasian and American tectonic plates in northern Iceland
Male Rock ptarmigan (Lagopus mutus) vocalising whilst in flight, showing winter plumage, Utsjoki, Finland, April
Water vole (Arvicola amphibius) standing on hind legs sniffing blackberry, Kent, UK, September
Bank vole (Clethrionomys glareolus) feeding on blackberries, Worcestershire, England, UK, August. Meetyourneighbours.net project
Red Irish lord (Hemilepidotus hemilepidotus) hides on the sebed, with Quillback rockfish (Sebastes maliger) and copper rockfish (Sebastes caurinus) behind in the bull kelp forest
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at sunrise. Kasanka National Park, Zambia
Sword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devil sSword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devils Trumpet Flower (Datura sp.). Yanacocha montane cloud forest, Ecuador
Deer mouse (Peromyscus maniculatus) on giant puffball mushroom, watching mosquito in the moonlight. Blackfoot Valley, Western Montana, USA, July
Gentoo Penguin (Pygoscelis papua) Wiencke Island, Antarctic Peninsula, Antarctica
Feral domestic rabbit (Oryctolagus cuniculus) juvenile running with dead leaf in mouth, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Tiger (Panthera tigris tigris), male portrait, Bandhavgarh, India, February 2013
Sandhill crane (Grus canadensis) landing at sunset. North America. Highly honoured in the Art in Nature category of the Natures Best Photography Windland Smith Rice International Awards Competition
European bison (Bison bonasus), Drawsko Military area, Western Pomerania, Poland, February
Lion (Panthera leo) cubs playing, Masai Mara Game Reserve, Kenya
Rock formations silhouetted at sunset on the Pacfic coast of Olympic National Park, Washington State, USA
Cowslips (Primula veris) growing on organic farm. Norfolk, UK, April
Wild Mustang pinto foal nuzzling up to mother, Sand Wash Basin Herd Area, Colorado, USA
Wild Mustang foals among wild flowers, Pryor Mountains, Montana, USA
Komodo Dragon (Varanus komodoensis) close up, Komodo National Park, Indonesia
White-tailed deer (Odocoileus virginianus) in snow, Acadia National Park, Maine, USA, February
British Longhair, kitten with blue-van colouration age 10 weeks in barn with straw
Mallard (Anas platyrhynchos) walking on ice, Acadia National Park, Maine, USA, April
Malayan tiger (Panthera tigris jacksoni), Malaysia. Captive. An Endangered species, only around 500 remain
Gannet (Morus bassanus) landing in colony, Bass Rock, Scotland, UK, July
Red fox (Vulpes vulpes) cub hiding in grass, Vosges, France, May
Graylag gosling (Anser anser) resting under parents wing, Pusztaszer, Hungary
Ice pile of broken shelf ice, near the shore of Lake Baikal, Siberia, Russia, March
Killer whale (Orcinus orca) tail slapping at sunset, transient race, Vancouver Island, British Columbia, Canada, July
Mountain Gorilla (Gorilla gorilla beringei) mother holding baby twins age five months, Parc National des Volcans, Rwanda. Endangered species