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Alpenrose (Rhododendron ferrugineum) in mountain landscape, Nordtirol, Austrian Alps June
Wild daffodil {Narciccus pseudonarcissus} flowering in hazel woodland, Peak District National Park, UK
Red deer (Cervus elaphus) stag in autumn woodlands, Bradgate Park, Leicestershire, UK, October
Red Squirrel (Sciurus vulgaris) in summer coat running across fallen log in heather, Scotland, UK. September
RF - European Badger (Meles meles) foraging at tree trunk in deciduous woodland. Mid Devon, UK. June
Emperor moth (Saturnia pavonia) caterpillar feeding on Ling heather (Calluna vulgaris) taken with fish eye view, Peak District National Park, Derbyshire, UK. August
Emperor moth (Saturnia pavonia) female wide angle view showing heather moorland habitat. Peak District National Park, UK. April
Brown hare (Lepus europaeus) adult running in wintery landscape. Derbyshire, UK, January
Red fox (Vulpes vulpes) at edge of stream, Kent, UK. January 2013
Red deer (Cervus elaphus) stag in heavy snowfall, Cairngorms National Park, Scotland, March 2012
European Badger (Meles meles) on the North Downs above Folkestone. Kent, UK, June. Camera trap photo
Mountain Hare (Lepus timidus) against mountains. Cairngorms National Park, Scotland, July
Male Adder (Vipera berus) tasting the air. Cannock Chase, Staffordshire, UK, October
Red Deer (Cervus elaphus) hind in early morning mist. Bradgate Park, Leicestershire, UK, September
Raven (Corvus corax) adult calling from rock in mountain habitat, Scotland, UK. November. Third place in British Birds Bird Photograph of the Year competition 2016
School of Longnose parrotfish (Hipposcarus harid) swarming over reef. Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Black rockfish (Sebastes melanops) in bull kelp (Nereocystis luetkeana), Browning Pass, Port Hardy, Vancouver Island, British Columbia, Canada
Klunzingers wrasse (Thalassoma klunzingeri) swimming over the edge of fringing reef where the water depth drops instantly from few feet to hundreds of meters. Ras Mohammed Marine Park, Sinai, Egypt
Pair of Sockeye salmon (Oncorhynchus nerka) on their redd in a shallow stream. Female in front of male. Huihil Creek, Adams River, British Colombia, Canada, October
Spring Crocus (Crocus vernus) in flower in snow, Campo Imperatore, Gran Sasso, Appennines, Abruzzo, Italy, May
Florida Red-bellied Turtle (Pseudemys nelsoni) Rainbow Springs, Rainbow Springs State Park, Florida, USA, March. Taken for the Freshwater Project
Grey heron (Ardea cinerea) beneath waterfall. Ambleside, Lake District, UK, November. Highly commended, Habitat category, British Wildlife Photography Awards (BWPA) competition 2012
Volcan Alcedo giant tortoises (Chelonoidis nigra vandenburghi) among steaming fumaroles on western caldera rim, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
African forest elephant (Loxodonta cyclotis) in water, Lekoli River, Republic of Congo (Congo-Brazzaville), Africa. Vulnerable species
White water lilies (Nymphaea alba) on surface of Danube delta rewilding area, Romania
Portrait of a wild Carpathian Grey Wolf (Canis lupus lupus) in snow-bound woodland habitat. Bieszczady, Carpathian Mountains, Poland, December
Fer-de-lance (Bothrops asper) camouflaged on the rainforest floor, Corcovado National Park, Osa Peninsula, Costa Rica
Parsons chameleon {Calumma parsonii} female walking along vine in tropical rainforest habitat. Andasibe-Mantadia NP, Madagascar
Large aerial pitchers of Veitchs pitcher plant (Nepenthes veitchii) growing up a tree trunk. Maliau Basin, Borneo
Morelets crocodile / Central American crocodile (Crocodylus moreletii) hiding in the plants at the surface of the cenote, Car Wash Cenote/ Aktun Ha, Yucatan peninsula, Mexico
Cheetah (Acinonyx jubatus) cubs 4 months, Masai-Mara Game Reserve, Kenya. Vulnerable species
Male Caucasian turs (Capra caucasica) rutting with Dykh-tau Mountain in the background, Kabardino-Balkarsky Zapovednik / Reserve, Russia, July
Silverback Mountain gorilla (Gorilla beringei) in the bamboo forest, this is Munyinya the leader of Hirwa group, Sabyinyo volcano, Volcano National Park, Rwanda
Crested tit (Lophophanes cristatus) clinging to lichen covered tree in snowfall, Scotland, UK February
Badger (Meles meles) foraging in woodland on edge of forest, Black Forest, The Black Forest, Germany, May. Highly commended in Mammals category, GDT competition 2011
Palm trees (Licuala valida) in swamp forest, Lambir NP, Borneo, Sarawak, Malaysia
Ecuadorian toad (Rhaebo ecuadorensis) on branch, Yasuni National Park, Orellana, Ecuador
Cup fungi (Cookeina sp) growing on decaying log, Danum Valley, Sabah, Borneo
Pitcher plant (Nepenthes veitchii x stenophylla), a natural hybrid. Maliau Basin, Borneo
Komodo Dragon (Varanus komodoensis) in habitat, Komodo National Park, Indonesia
Mountain gorillas (Gorilla beringei) Hirwa group led by the silverback dominant male Munyinya, Volcanoes National Park, Rwanda, elevation 2610 m
Quiver tree forest (Aloe dichotoma) at night with stars and the Milky Way, Keetmanshoop, Namibia
Close up of colourful mixed woodland in autumn, Kinnoull Hill Woodland Park, Perthshire, Scotland November 2011
A misty morning over a mixed woodland in autumn. Kinnoull Hill Woodland Park, Perthshire, Scotland, November 2011
Sunset over savanna landscape image with a lone (Acacia) tree, Masai Mara NR, Kenya
Footpath through Wild Garlic / Ramsons (Allium ursinum) carpeting deciduous woodland, North Somerset, UK
Zebra (Equus quagga) foal running through Devils Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January
Crowned dwarf / Smooth-fronted caiman (Paleosuchus trigonatus) with mouth open, Cuyabeno, Sucumbios, Ecuador