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A female Spiny Seahorse (Hippocampus guttulatus) in weeds. Sardina, Gran Canaria, Canary Islands, East Atlantic Ocean, October
A male Weedy Seadragon (Phyllopteryx taeniolatus) carrying eggs along tail swims through a Kelp (Macrocystis pyrifera) forest in Tasmania, Australia
A misty morning view over a mixed woodland in autumn, Kinnoull Hill Woodland Park, Perthshire, Scotland, November 2011
Thomsons gazelle (Eudorcas thomsonii) herd with Wildebeest herd visible beyond, Masai-Mara game reserve, Kenya
Black tailed prairie dogs on burrow mound {Cynomys ludocicianus}, Badlands, USA
Fallow deer (Dama dama) amongst bracken in oak woodland, Cheshire, UK, August, Highly Commended, Habitat category, British Wildlife Phototgraphy Awards (BWPA) competition 2011
Marine iguana (Amblyrhynchus cristatus) on coast, Galapagos
African elephants (Loxodonta africana) large family group on migration, Amboseli National Park, Kenya
American flamingo (Phoenicopterus ruber) feeding in water with hill in background. Floreana Island, Galapagos, Ecuador, June
Black bear (Ursus americana) foraging for alpine berries during Autumn, on hillside, Cascade Mountains, Washington, USA
Guanaco (Lama guanicoe) herd below Torres del Paine, Torres del Paine National Park, Patagonia, Chile
Chamois (Rupicapra rupicapra) running through snowy forest. Vosges mountain, France, November
White-tailed Deer (Odocoileus virginianus) in front of forest, with reflection of trees in water, Acadia National Park, Maine, USA, October
Bittern (Botaurus stellaris) camouflaged as it stands motionless in reedbed. Yorkshire, UK, January (non-ex)
Willow grouse (Lagopus lagopus) camouflaged in snow. Utsjoki, Finland. April 2007
Great Grey Owl (Strix nebulosa) chicks in nest, Sweden
Leopard (Panthera pardus) male climbing on fallen tree, , Nakuru national park, Kenya
Herd of Southern giraffes (Giraffa camelopardalis giraffa) and Burchells zebras (Equus quagga) aerial view, Okavango delta, Bostwana
Mountain Clouded Yellow (Colias phicomone) profile portrait, Aosta Valley, Monte Rosa Massif, Pennine Alps, Italy
Aerial view over subtropical mangrove wetlands of the Everglades National Park. Florida, USA, February 2012
Gulik Falls, edge of southern plateau, Maliau Basin, Sabahs Lost World, Borneo (digitally stitched image)
Malachite kingfisher (Alcedo cristata) perched on rush, Intaka Island, Cape Town, South Africa, February
Brown trout (Salmo trutta) resting on river bed near the Aigas Field Studies Centre, Inverness-shire, Scotland, July
Spotted owl (Strix nebulosa) Gifford-Pinchot National Forest, Washington USA
Bridled nailtail wallaby (Onychogalea fraenata) Queensland, Australia, Endangered species, October
Dipper (Cinclus cinclus) perched on moss-covered waterfall, Peak District NP, Derbyshire, UK, May, Highly Commended, Habitat category, British Wildlife Phototgraphy Awards (BWPA) competition 2011
Black Rhinoceros (Diceros bicornis) charging. Etosha National Park, Namibia, January
Red Deer (Cervus elaphus) hinds silhouetted in grassland. Richmond Park, London, UK, October
Pheasant (Phasianus colchicus) male amongst Bluebell flowers, UK, April
A male Leafy Seadragon (Phycodurus eques) swimming. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
A Pygmy Seahorse (Hippocampus bargibanti) camouflaged in red Seafan (Muricella sp.). Pygmy Seahorses are small, most less than 15mm in length. Tulamben, Bali, Indonesia, October
A juvenile Weedy Seadragon (Phyllopteryx taeniolatus) in its habitat. Flinders Jetty, Melbourne, Victoria, Australia, March
Red Fox (Vulpes vulpes) resting on straw bale in field. Vosges, France, July
Eurasian Sparrowhawk (Accipiter nisus) rear view perching on fence post with tail feathers spread. Vosges, France, January
Black-legged Kittiwake (Rissa tridactyla) on ice. Floe edge, Arctic Bay, Nunavut, Canada, April
Polar Bear (Ursus maritimus) walking on icepack with arctic coast in the background. Floe edge, Arctic Bay, Baffin Island, Nunavut, Canada, April
Konik Wild Horse (Equus ferus caballus) in grasslands with Purple loosestrife flowers (Lythrum salicaria) in foreground. The Netherlands, July
Reef with Pygmy Sweepers (Parapriacanthus ransonetti), Barrel Sponge (Xestospongia testudinaria), and soft corals. Komodo National Park, Indonesia, October
Red deer (Cervus elaphus) stag amongst winter vegetation, Salburua Park, Alava, Spain, November
Catesbys snail-eating snake (Dipsas catesbyi) on Heliconia plant, Pacaya Samiria National Reserve, Amazon Basin, Loreto, Peru
Golden eagle (Aquila chrysaetos) in flight over snow, Alps, Austria, controlled conditions
Panther Chameleon {Furcifer pardalis} walking along branch on coast, Masoala Peninsula National Park, north east Madagascar
Caribbean reef shark (Carcharhinus perezi) swims over a coral reef. Grand Bahama Island, Bahamas. Tropical West Atlantic Ocean
Crested Newt (Triturus cristatus carnifex) underwater view of female swimming on surface, Traversari lake, Campigna National Park, Emilia Romagna and Tuscany regions, Italy
Flooded birch and alder woodland in autumn, Cairngorms National Park, Scotland, UK, November, RUNNER UP: In Praise of Plants category of the Wildlife Photographer of the Year Competition 2010
Lammergeier (Gypaetus barbatus) flying over the Pyrenees mountains, Aragon, Spain
Beech Woodland {Fagus sylvatica} at sunrise, late summer, Mendip Hills AONB, Somerset, UK, 2008
Desert scorpion {Parabuthus villosus} Namib desert, Namibia
