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Great Grey Owl (Strix nebulosa) in flight, with forest behind, Rovaniemi Finland March
Blue Sheep (Pseudois nayaur) on hillside, Kekexili, Qinghai, China, January
Wolves (Canis lupus) sitting on a hillside in snow. Yellowstone, USA, February
Grey wolf (Canis lupus) Katmai National Park, Alaska, USA, August
Wide angle close-up of two European grey wolves (Canis lupus), captive, Norway, February
European grey wolf (Canis lupus) in landscape, captive, Norway, February
Wide angle close-up of a European grey wolf (Canis lupus) in landscape, captive, Norway, February
Speckled Wood (Pararge aegeria) male flying in habitat, Finland, April
Red foxes (Vulpes vulpes) fighting in snowy field with woodland in the background, Vosges, France, February
Rainbow trout (Oncorhynchus mykiss) under trees, Capernwray Lake, Lake District, England, UK
Polar bear (Ursus maritimus) walking in snow, Svalbard, Norway, April
Brown bear (Ursus arctos) in autumnal forest, Finland, September
Bison (Bison bison) herd walking in line in snow, Yellowstone National Park, Wyoming, USA, February
Moose Bull (Alces alces) walking in forest clearing, Denali National Park, USA, September
Frigga Fritillary (Boloria frigga) in habitat over cotton grass (Eriophorum) Finland, June
Musk Oxen (Ovibos moschatus) in mountain habitat. Dovrefjell National Park, Norway, September
Musk Ox (Ovibos moschatus) in mountainous habitat. Dovrefjell National Park, Norway, September
Brown Bear (Ursus arctos) and reflection in pond. Finland, Europe, June
Brown Bear (Ursus arctos) by water, reflected. Finland, Europe, June
Cobalt-winged parakeets (Brotogeris cyanoptera) eating clay. Yasuni National Park, Orellana, Ecuador
Sunset above the canopy in the Peruvian Amazon. Tambopata Reserve, Madre de Dios, Peru. August, 2016
RF - Snow leopard (Panthera uncia) male moving over rocky, snow-covered slopes. Ulley Valley, Himalayas, Ladakh, India
Snowy owl (Bubo scandiaca) female sitting among snow covered vegetation on ground, Canada February
Dense rainforest surrounding estuary mouth, Corcovado National Park, Osa Peninsula, Costa Rica
Trumpet / Stemless Gentian (Gentiana acaulis) Nordtirol, Austrian Alps. June
Fallow deer heard (Dama dama) in heavy snow. Chatsworth Park, Derbyshire, Peak District National Park. January
Viscid primrose (Primula latitolia) flowering in snow. Nordtirol, Austrian Alps, July
Alpine Aster (Aster alpinus), Aosta Valley, Monte Rosa Massif, Pennine Alps, Italy. July
Wild garlic / Ramsons / Bear Garlic (Allium ursinum) and Bluebells (Endymion nonscriptus) flowering in deciduous woodland. Peak District National Park, Derbyshire, UK, May, 2009
Viscid primrose (Primula latitolia) on mountainside, Nordtirol, Austrian Alps, 2600 metres, June
Dwarf snowbell (Soldanella pusilla) in flower on mountainside, Austrian Alps, June
Alpine moon daisy (Leucanthemopsis alpina) on mountainside, fisheye lens. Nordtirol, Austrian Alps, Austria, June
Red Deer Stag (Cervus elaphus) amongst Autumn vegetation, Bradgate Park, Leicestershire, UK, October 2015
Red squirrel (Sciurus vulgaris) on snow-covered branch in pine forest, Glenfeshie Estate, Cairngorms NP, Highlands, Scotland, UK, January
Kittiwake (Rissa tridactyla) on nest built on cliff, with long exposure of sea below in the background, Isle of Lunga, Treshnish Isles, Scotland, June
Black Grouse (Tetrao tetrix) males at spring lek. Deeside, Scotland, April
Short eared owl hunting by day {Asio flammeus} Scotland
Capercaillie (Tetrao urogallus) male displaying in pine forest in snow, Cairngorms NP, Highlands, Scotland, UK, January. Photographer quote
A long exposure renders Bar jacks (Carangoides ruber) as ghostly trails as they swim around a sponge, on a coral reef, at night. East End, Grand Cayman, Cayman Islands
Group of Circular spadefish (Platax orbicularis) gather at a cleaning station beneath a table coral (Acropora sp.) on a coral reef. Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea
Dipper (Cinclus cinclus) perched on moss covered waterfall, Derbyshire, UK
A female Spiny Seahorse (Hippocampus guttulatus) in weeds. Sardina, Gran Canaria, Canary Islands, East Atlantic Ocean, October
A male Weedy Seadragon (Phyllopteryx taeniolatus) carrying eggs along tail swims through a Kelp (Macrocystis pyrifera) forest in Tasmania, Australia
A misty morning view over a mixed woodland in autumn, Kinnoull Hill Woodland Park, Perthshire, Scotland, November 2011
Thomsons gazelle (Eudorcas thomsonii) herd with Wildebeest herd visible beyond, Masai-Mara game reserve, Kenya
Black tailed prairie dogs on burrow mound {Cynomys ludocicianus}, Badlands, USA
Fallow deer (Dama dama) amongst bracken in oak woodland, Cheshire, UK, August, Highly Commended, Habitat category, British Wildlife Phototgraphy Awards (BWPA) competition 2011
Marine iguana (Amblyrhynchus cristatus) on coast, Galapagos