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Raven (Corvus corax) adult calling from rock in mountain habitat, Scotland, UK. November. Third place in British Birds Bird Photograph of the Year competition 2016
Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Carp (Cyprinus carpio) Aare River, Switzerland 2006
Robin (Erithacus rubecula) at RSPB Leighton Moss Bird Reserve, Silverdale, Lancashire, England, UK. February 2016
Underwater river scene with freshwater plants and Tetra fish, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul
Atlantic Puffin (Fratercula artica) adult on breeding cliffs. Hermaness NNR, Shetland, UK, June
Piraputanga fish (Brycon hilarii) in underwater landscape, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul, Brazil
RF - Southern stingray (Dasyatis americana) swimming over sand in shallow water at dawn, Cayman Islands, Caribbean Sea
Emperor penguin (Aptenodytes forsteri) viewed through hole in iceberg at Snow Hill Island rookery, Antarctica. November
Asian Elephant (Elephas maximus) male, India
Cuckoo flower or Ladys smock (Cardamine pratensis) on Hawkesbury Common, Gloucestershire, England, UK. April
Coral reef scenery with Gorgonian and shoal of Pygmy / Glassy sweepers (Parapriacanthus guentheri) Egypt, Red Sea
Asian Elephant (Elephas maximus) male, walking through forest, South India
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Wild daffodil (Narcissus pseudonarcissus) Bay, Ullswater, Lake District, England, UK, March
Scottish primrose (Primula scotica) native only on the northern coast of Scotland and Orkney, here growing on grasslands at Dunnet Head, Caithness, England, UK, July
Madagascar pitcher plants {Nepenthes madagascariensis} Pangalanes canal, East Madagascar
Red fox (Vulpes vulpes) at night in snow, camera trap image, Jura Mountains, Switzerland, August. Commended in the Mammals category of the Asferico Competition 2016
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
Marine iguana (Amblyrhynchus cristatus) on shore, Punta Moreno, Isabela Island, Galapagos
Whooper swan (Cygnus cygnus) in landscape of Landmannalaugar, Iceland, June
Male Black rhinoceros (Diceros bicornis) running, Nakuru National Park, Kenya
Herd of Tibetan Wild Ass (Equus kiang) with view of snow capped mountains behind, Tso Kar lake, Ladakh, India, June 2010
Caribou / Reindeer bull (Rangifer tarandus) close-up grazing, Denali National Park, Alaska, USA, September
RF- Mute swan (Cygnus olor) on water in winter dawn mist. Loch Insh, Cairngorms National Park, Highlands, Scotland UK, December
Glen Affric in autumn, Highlands, Scotland, UK, October 2012
King cobra (Ophiophagus hannah) in strike pose, Malaysia
Arctic Fox (Alopex / Vulpes lagopus) standing on ridge, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
A male Leafy Seadragon (Phycodurus eques) carrying eggs. Wool Bay, Edithburgh, South Australia, Gulf of St Vincent, November
English bluebell (Hyacinthoides non-scripta) Lower Woods, Gloucestershire, England, UK, April
Common toad (Bufo bufo) tadpole swimming seen from below, Belgium, June
Blue poppy (Meconopsis zangnanensis) Mt Qomolangma National Park, Qinghai Tibet Plateau, China
Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement
Alpine ibex (Capra ibex) female, Jura, Switzerland, September
Florida manatee (Trichechus manatus latirostris) swimming into a fresh water spring, Three Sisters Spring, Crystal River, Florida, USA
The forests of the Modi Khola river valley shrouded in fog. Annapurna Sanctuary, central Nepal, November 2011
Timber Wolves (Canis lupus) resting on a rock by the shoreline. Near Mussel Inlet, Great Bear Rainforest, British Columbia, Canada, October
Water lilies (Nymphaeaceae) seen from below, Cenote Car Wash / Aktun Ha, Mexico. Yucatan peninsula
Water Vole (Arvicola amphibius / terrestris) foraging by water. Kent, UK, August
Piraputanga (Brycon hilarii) shoal, Rio Olhio d agua, tributary of Rio da Prata, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil
Alpine marmot (Marmota marmota), with Mount Grossglockner (3798m) in background, Hohe Tauern National Park, Austria, July
Edwards Atlas Moth (Attacus edwardsii) in defensive posture, Bhutan, June
Emergent Menggaris Tree / Tualang (Koompassia excelsa) protruding the canopy of lowland rainforest. Danum Valley, Sabah, Borneo
White stork (Ciconia ciconia) in tree, migrating towards Europe, Masai-Mara Game Reserve, Kenya
African elephant (Loxodonta africana) herd walking in the plains during the dry season, Masai-Mara Game Reserve, Kenya. October
Giant pacific octopus (Enteroctopus dofleini) swimming freely after release from captivity, Vancouver Island, British Columbia, Canada, Pacific Ocean
Red Fox (Vulpes vulpes) walking along a fallen trunk in misty forest. Black Forest, Germany, November
Species rich alpine meadow with Orange lily (Lilium bulbiferum), Meadow clary (Salvia pratensis) and Yellow rattle (Rhinathus sp)
Red-and-Green Macaws (Ara chloropterus), feeding at the wall of a riverside clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November
Common blue butterfly (Polyommatus icarus) resting on Devil's bit scabious (Succisa pratensis), Volehouse Farm, Devon, UK. August
Sanderling (Calidris alba) flock foraging at sunset, Britanny, France, October
Peregrine Falcon (Falco peregrinus) perched on a cliff above a woodland in autumn. Kinnoull Hill Woodland Park, Perthshire, Scotland, November
Common loon (Gavia immer) nesting at edge of lake and calling, Acadia National Park, Maine, USA, August
Alpine ibex (Capra ibex), male in habitat, Bernese Alps, Switzerland. July
Scots pine trees (Pinus sylvestris) in mist at sunrise, Beinn Eighe NNR, Torridon, NW Scotland, UK, June
Red Fox (Vulpes vulpes) in distance in snow habitat. Vosges, France, December
Boomslang (Dispholidus typus) Immature male hunting in thorn bush above deHoop Vlei, Western Cape, South Africa, December
Eastern Grey Bamboo Lemur (Hapalemur griseus) feeding on bamboo shoot, Andasibe-Mantadia National Park, Madagascar, October
Healthy coral reef landscape, Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea
Great blue heron (Ardea herodias) taking off, with coastal landscape, Las Bachas, Santa Cruz Island, Galapagos
Young Badger (Meles meles) foraging in woodland on edge of woodland, The Black Forest, Germany, May
Landscape of Zanskar River, a tributary of the Indus, borders part of Hemis national park, habitat of the snow leopard (Panthera uncia), Hemis National Park, Ladakh, India
Grizzly bear (Ursus arctos horribilis) and Grey wolf (Canis lupus) on beach, Katmai National Park, Alaska, USA, August
Sunrise over coastal mudflats, Campfield Marsh RSPB reserve, Bowness, Solway Firth, Cumbria, UK, April 2011
Bison (Bison bison) female with calf walking through snow in front of frost-covered forest, Yellowstone National Park, USA
Mixed flock of Snow geese (Chen caerulescens atlanticus / Chen caerulescens) and Sandhill cranes (Grus canadensis) at wintering area at dawn, with full moon, Bosque del Apache, New Mexico, USA
Common spotted orchid (Dactylorhiza fuchsii) on a roadside verge near Bristol, England, UK, June
Alpine aster (Aster alpinus) flowering in alpine meadow, Alps, Engadine, Switzerland. July
Atlas beetle / Three-horned rhinoceros beetle male (Chalcosoma sp.) with enourmous horns, used to fight rival males. Danum Valley, Sabah, Borneo
Tajinaste rojo (Echium wildpretii), Teide National Park, UNESCO World Heritage Site. Canary Islands. Endemic
Western hemlock (Tsuga heterophylla) in Douglas-fir forest (Pseudotsuga menziesii), Oregon, USA, June
Tibetan gazelle / Goa (Procapra picticaudata) Keke Xili / Hoh Xil Nature Reserve, Tibetan High plateau, Qinghai, China
Lemon shark juvenile (Negaprion brevirostris) trying to feed on the leaves of a red mangrove (Rhizophora mangle). As pups, the sharks have to learn to hunt for themselves and sometimes make mistakes
Fire salamander (Salamandra salamandra) in forest habitat, Hallerbos, Belgium. May
Central bearded dragon (Pogona vitticeps) male on stick in chenopod shrubland habitat, Meringur in NW Victoria, Australia
White-lipped snail (Cepaea hortensis) crawling over an oak sleeper retaining a garden lawn with a greenhouse in the background, Wiltshire, UK, April
Field of Paramo flower / Frailejones (Espeletia pycnophylla), highland paramo, northern Ecuador
Umbrella thorn tree (Vachellia tortilis) in the Etosha pan with common ostrich (Struthio camelus) male. Etosha National Park, Namibia
Oak tree (Quercus robur) in winter mist, parkland, Herefordshire Plateau, England, February
Yellow whitlow grass (Draba aizoides) growing at Pennard Castle, South Wales, UK. March. In Britain into only occurs on the cliffs around the Gower Peninsula
Wasp orchid, (Ophrys apifera var. trollii) growing on Cleeve Hill near Cheltenham, Gloucestershire, England, UK, July
Rosy garlic (Allium roseum) naturalised in the Avon Gorge, Bristol, England, UK, June