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Northern wheatear (Oenanthe oenanthe) adult male in spring plumage feeding on dung flies at farm midden heap, Hertfordshire, UK, April
Flock of Barnacle geese (Branta leucopsis) feeding on grazing marshes, Caerlaverock WWT, Dumfries and Galloway, Scotland, UK, December
Large flock of Barnacle geese (Branta leucopsis) taking off from grazing marshes, Caerlaverock WWT, Dumfries and Galloway, Scotland, UK, December
Common seals (Phoca vitulina) hauled out on saltmarsh, Wallasea Wild Coast Project, RSPB Greater Thames Futurescapes Project, Wallasea Island, Essex, England, UK, October
Four Red deer stags (Cervus elaphus) walking and grazing on open snow-covered moorland, near woodland edge, Cairngorms NP, Scotland, UK, December
Arctic charr (Salvelinus alpinus) males showing breeding colours, in a river where they have come to spawn, Ennerdale Valley, Lake District NP, Cumbria, England, UK, November 2011
Five Brown hares (Lepus europaeus) silhouetted against the skyline, Hertfordshire, UK, March 2010
Bed of Common mussels (Mytilus edulis) growing on rocks, feeding, Megavissey, Cornwall, UK, May
Konik horse (Equus caballus) herd grazing with two stallions interacting, Wicken Fen, Cambridgeshire, UK, June 2011
Common / Eurasian cranes (Grus grus) three 18 month juveniles, Bart, Christine and Gilbert, released by the Great Crane Project onto the Somerset Levels and Moors
Flamingos on Laguna / Lake Colorado at dawn, Eduardo Avaroa Andean Fauna National Reserve, Bolivia, December 2009
Turkmenian Wild Ass / Kulan (Equus hemionus kulan) Badkhyz, Turkmenistan. Endangered
African elephant (Loxodonta africana) herd walking in procession, Kaokoveld Desert, Namibia
Walia ibex (Capra walie) herd, Simien Mountains National Park, Ethiopia
Japanese fireflies (Luciola cruciata) in flight at night, endemic species, Yaku-shima UNESCO World Heritage Site, Kagoshima, Japan, May
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Close-up of group of toadstools (Mycena sp) autumn. Leicestershire, UK. November. Photographer quote: It is all too easy to overlook the fascinating miniture world underneath our feet - mosses
Grey wolves playing in snow {Canis lupus} captive, USA
American bald eagle (Haliaeetus leucocephalus) three perched, Homer, Alaska, Jan
Cows (Bos taurus) in field of buttercups looking over dry stone wall. Peak District National Park, Derbyshire, UK, May 2012
Sika deer (Cervus nippon), stag, hind and young, amongst flowering heather, Arne RSPB reserve, Dorset, England, UK, August
Common starling (Sturnus vulgaris) flock at dusk engaging in pre-roost display, Dumfries-shire, Scotland, March
Pasque flowers (Pulsatilla vulgaris) in rain, Lorraine, France, April. 3rd Prize in the Other Nature Topics category of Melvita Nature Images Awards 2013
Common / Eurasian crane (Grus grus) flock in flight with snow topped Pyrenees mountains in the background, during southward migration period, Haute Pyrenees, Gascony, France. December 2009
Wild flower meadow, Common poppy (Papaver rhoeas) Derbyshire, UK
Boer domestic goat (Capra hircus) waiting to be fed, Norfolk, UK, September
Elk / Wapiti (Cervus canadensis) herd with young on meadowland against the Grand Tetons, low clouds and the Snake River. Grand Teton National Park, Wyoming, USA, June
Massing herds of White bearded wildebeest (Connochaetes taurinus albojubatus) on migration, Serengeti National Park, Tanzania
Kittiwake (Rissa tridactyla) flock feeding in from of the Monaco Glacier, Leifdefjord, Svalbard, Norway 2010
Japanese macaque / Snow monkey {Macaca fuscata} young monkeys playing in the sunshine in the trees, Jigokudani, Nagano, Japan
Wildebeest (Connochaetes taurinus) crossing the Mara river, Masai-Mara Game Reserve, Kenya
American Buffalo / Bison (Bison bison) grazing in open plains. Yellowstone National Park, Wyoming, October
A group of meerkats (Suricata suricatta) or suricates, stand on an old termite mound to gain extra height and watch for predators in their territory on the edge of Makgadikgadi Pans National Park
Tibetan antelope (Pantholops hodgsonii) herd in the early morning with a low moon, Kekexili, Qinghai, Tibetan plateau, China, January
Cornflower (Centaurea cyanus) UK, July
Alpine ibex (Capra ibex) adult male chasing two young males away, Gran Paradiso National Park, Italy, July
Domestic cows (Bos taurus) and free roaming horses grazing in the Pyrenees-Atlantiques, Pyrenees, France, June
Rothschilds giraffe (Giraffa camelopardalis rothschildi), Murchison Falls National Park, Uganda, East Africa. Murchison Falls National Park hosts the only remaining naturally occurring population of
Blacktip reef sharks (Carcharhinus melanopterus) swimming in shallow crystal clear water, Aldabra Atoll, Seychelles, Indian Ocean
Ebdem Geese Goslings at four days
A flock of Greylag Goose (Anser anser) flying in formation above misty fields. The Netherlands, March
Tench (Tinca tinca) on the surface of a turbid marsh pond, with Rudd (Scardinius erythrophthalmus) above, Fribourg, Switzerland, 2004
Gulls on the beach at sunset, The Wash, Hunstanton, Norfolk, UK, September 2011
Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October
Little Auks (Alle alle) flying low above surface in front of iceberg. Spitsbergen, Svalbard, Norway, July 2010. Highly commended in the Birds category in the GDT competition 2011
Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December
King penguin (Aptenodytes patagonicus) chicks in mass breeding colony, St Andrews Bay, South Georgia. Taken on location for BBC Frozen Planet series, 2008
King penguins (Aptenodytes patagonicus) on beach, South Georgia. Taken on location for BBC Frozen Planet series, 2008