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Badger (Meles meles) two outside sett interacting, Vosges, France, May
Mute swan (Cygnus olor) pair preening in unison, Gloucestershire, UK, December
Swallow-tailed gull (Creagrus furcatus) pair mutual preening in courtship. Genovesa Island, Galapagos Islands, Ecuador, June
American flamingo (Phoenicopterus ruber) preening feathers. Captive
King penguin (Aptenodytes patagonicus), two sleeping on snow. Grytviken, South Georgia. March
African Spoonbill (Platalea alba) preening, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Snares Crested penguin (Eudyptes robustus) bathing in freshwater pool, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species
Black-browed albatross pair (Thalassarche melanophris) mutual grooming during copulation. New Island, Falkland Islands, November
European river otter (Lutra lutra) grooming, Shetland, Scotland, UK, September
Tabby kitten grooming paw
Royal spoonbill (Platalea regia) preening and shaking head feathers, Victoria, Australia
Impala (Aepyceros melampus) with redbilled oxpecker (Buphagus erythrorhynchus), Kruger National Park, South Africa
European brown bear (Ursus arctos) scratching up against tree in snow, Finland, April
Leopard (Panthera pardus) female grooming cub age 6 months Little Kwara, Botswana June
Egytian Goose (Alopochen aegyptiaca) rear view bathing, sings spread, Zimanga Private Game Reserves, South Africa
Jaguar (Panthera onca) four month old cub washing, native to Southern and Central America, captive
Cheetah (Acinonyx jubatus) mother grooming cub, Masai-Mara Game Reserve, Kenya. Vulnerable species
Gyrfalcon (Falco rusticolus) preening, Myvatn, Thingeyjarsyslur, Iceland, June 2009
Red footed falcon (Falco vespertinus) on branch preening, at sunset, Hortobagy National Park, Hungary, May 2009
Two Konik horse foals mutual grooming, Oostvaardersplassen, Netherlands, June 2009 WWE BOOK
Ruddy shelduck (Tadorna ferruginea) preening, Bagerova Steppe, Kerch Peninsula, Crimea, Ukraine, July 2009
White pelicans (Pelecanus onocrotalus) preening, Lake Belau, Moldova, June
Grey seal (Halichoerus grypus) with flipper in its mouth, Donna Nook, Lincolnshire, UK, November 2008
Grey seal (Halichoerus grypus) pup with flipper in its mouth, Donna Nook, Lincolnshire, UK, November 2008
Konik horse (Equus caballus) mutual grooming, wild herd in rewilding project, Wicken Fen, Cambridgeshire, UK, June 2011
Pair of Gannets (Morus bassanus) mutual preening, Bass Rock, Firth of Forth, Scotland, UK, June
Mute swan (Cygnus olor) juvenile on water preening, Fife, Scotland, UK, November
Erect-crested penguins (Eudyptes sclateri) bathing and preening prior to moulting. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
Japanese Macaques (Macaca fuscatata) grooming in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Florida manatee (Trichechus manatus latirostrus) being cleaned by blue gill sunfish (Lepomis macrochirus). Three Sisters Spring, Crystal River, Florida. USA. February 2010
Little egret (Egretta garzetta) shaking water off wings at sunset, Lake Csaj, Pusztaszer, Hungary. May. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Wild horse / mustang, mare rolling, Pryor Mountains, Montana, USA, July