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Two African scops owls (Otus senegalensis) perched in tree, Katavi National Park, Tanzania
Ibex (Capra ibex) feeding on grass, head portrait, Schwyz, Switzerland. August
Barn owl (Tyto alba) perched at entrance to tree hollow, Hampshire, UK. October. Captive, controlled conditions
Arctic fox (Vulpes lagopus) resting in grass, Isfjorden, Svalbard, Norway, July
Vestrahorn mountain at dawn, south-east Iceland, Atlantic Ocean. November
Small tortoiseshell butterfly (Aglais urticae) basking on Cornish slate wall, Crantock, Cornwall, UK. June
Red rock crab (Grapsus adscensionis) grazing on algae from rock, Tenerife, Canary Islands, October
Kea (Nestor notabilis) juvenile, taking flight off rock, Arthur's Pass National Park, South Island, New Zealand
Dapple Dachshund puppy with Netherland dwarf rabbit and grey Tabby kitten sitting side by side, portrait
Swallow-tailed gull (Creagrus furcatus) eye detail, San Cristobal Island, Galapagos National Park, Galapagos Islands
Santa Catalina Island rattlesnake (Crotalus catalinensis), only rattlesnake without rattle, slithering in rock. Santa Catalina Island, Loreto Bay National Park, Sea of Cortez, Mexico. May
Female Siberian flying squirrel (Pteromys volans orii) sitting at entrance to nest. Hokkaido, Japan. February
Male Siberian flying squirrel (Pteromys volans orii) attempting to enter nest with female inside during reproductive season but stopped mid-way then rejected. Hokkaido, Japan. March
Composite image showing Siberian flying squirrel (Pteromys volans orii) defecating for duration of about twenty seconds after emerging from nest to forage. Hokkaido, Japan. March
Split-second prior to Siberian flying squirrel (Pteromys volans orii) landing on tree, adjusting its attitude in mid-flight toward relatively vertical position to reduce speed
Pair of Lava gulls (Leucophaeus fuliginosus) during courtship, Genovesa Island, Galapagos Islands, Ecuador
Pinkfooted goose (Anser brachyrhynchus) on burnt heather moorland, Berwickshire, Scottish Borders, Scotland, UK. October
Bay of Laig, Isle of Eigg towards Isle of Rum, Inner Hebrides, Scotland, UK, April 2014
Lava gull (Larus fuliginosus) on nest, Mosquera Islet, Galapagos, South America
Young, female Grey seal (Halichoerus grypus) swimming over a forest of Cuvie kelp (Laminaria hyperborea), Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Spoonbill (Platalea leucorodia) calling in shallow pond on misty morning, Valkenhorst Nature Reserve, The Netherlands, August
Mixed group of Grey heron (Ardea cinerea), Great white egret (Ardea alba) and Spoonbill (Platalea leucorodia) on misty morning in shallow pond, Valkenhorst Nature Reserve, The Netherlands, August
Two adult Tufted titmice (Baeolophus bicolor) surrounded by snow, winter, New York, USA
Sanderling (Calidris alba) Britanny, France, October
Stormy sky over Klein Schietveld, Brasschaat, Belgium. March 2019
Women walking the dog in heavy snow near Ambleside, England, UK. March 2006
Espanola mockingbird (Mimus macdonaldi) on beach, Galapagos
Silver tabby kitten, Freya, age 7 weeks, with raised paws
Shaggy grey cat, portrait
Head portrait of grey Andalusian stallion running with mane flowing in arena in Northern France, Europe
Blue-and-white and black kittens
Puss moth (Cerura vinula) near Lough Bunny, County Clare, Ireland. April. Small repro only
Lichen on rocks, Assynt Uplands, Scotland, UK, January
Flock of juvenile Common / Eurasian cranes (Grus grus) recently released by the Great Crane Project onto the Somerset Levels and Moors, fly over grazing cattle. Somerset, UK, October 2010
Juvenile Common / Eurasian cranes (Grus grus) recently released by the Great Crane Project onto the Somerset Levels and Moors
Greylag goose (Anser anser) flying, Glenfeshie, Cairngorms National Park, Scotland, UK, April
Blue-footed booby (Sula nebouxii) plunge-diving at high speed, San Cristobal Island, Galapagos, Ecuador
Lioness (Panthera leo) and Thomsons gazelles (Eudorcas thomsonii) silhouetted at sunset, Masai-Mara game reserve, Kenya
Young Baikal seal (Pusa sibirica) at breathing hole. Lake Baikal, Russia, April
Salvins albatross (Thalassarche salvini) flying at sea, closeup of head. Kaikoura, Canterbury, New Zealand, September. Vulnerable species
Male Long tailed duck (Clangula hyemalis) on water, Batsfjord, Norway, March
Two male Bearded reedling (Panurus biarmicus) eating seeds from the spike of a common bulrush (Typha latifolia) in a reedbed, Belgium, January. Digital composite
Phayers Leaf monkey (Trachypithecus phayrei) Gaoligongshan NP, Yunnan province, China
Dendritic drainage patterns eroded into sand. Isle of Coll, Scotland, October 2012
Robin (Erithacus rubecula) in snow. Dorset, UK, January 2010
Blackcap (Sylvia atricapilla) Uto Finland May
Humpback Whale (Megaptera novaeangliae) female. Tonga, South Pacific, September
Blue Sheep (Pseudois nayaur) on hillside, Kekexili, Qinghai, China, January