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Feral kitten with sleeping Galapagos sea lion (Zalophus wollebaeki) Puerto Fragata, Isabela Island, Galapagos
Espanola saddelback tortoise (Chelonoidis hoodensis) pair mating, Espanola Island, Galapagos
Vegetarian finch (Platyspiza crassirostris) snips petals from a hardy vine, Fernandina Island, Galapagos
Short-eared owl (Asio flammeus) in flight, Santa Fe Island, Galapagos
Galapagos penguin (Spheniscus mendiculus) and Marine iguana (Amblyrhynchus cristatus) male, Isla Tortuga, Isabela Island, Galapagos
Blue-footed booby (Sula nebouxii) with chicks begging for food, Santa Cruz Island, Galapagos, Ecuador
Pink iguana (Conolophus marthae) Northern caldera rim, Wolf Volcano, Isabela Island, Galapagos Islands, Ecuador, Critically endangered species
Panamic Cushion Sea Star (Pentaceraster cumingi) and a school of Yellow-tailed Grunt (Anisotremus interruptus), Galapagos Island, Ecuador. December
Magnificent frigatebird (Fregata magnificens) in flight over sea between Isabela Island and Fernandina Island, Galapagos
Galapagos sea lion (Zalophus wollebaeki) jumping out of Pacific ocean. Sombrero Chino, Santiago Island, Galapagos
Scalloped hammerhead shark (Sphyrna lewini) swims through a school of Pacific creolefish (Paranthias colonus). Wolf Island, Galapagos National Park, Galapagos Islands.. East Pacific Ocean
Marine Iguanas (Amblyrhynchus cristatus) basking. Fernandina Island, Galapagos Islands
Red-footed booby (Sula sula) perched in tree. Genovesa Island, Galapagos
Nazca booby (Sula granti), pair on rocky shore at dawn. Punta Suarez, Espanola Island, Galapagos. May 2015
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Marine iguana (Amblyrhynchus cristatus), Punta Vicente Roca, Isabela Island, Galapagos
American flamingo (Phoenicopterus ruber) group of four with one displaying Punta Moreno, Isabela Island, Galapagos
Swallow-tailed gull (Creagrus furcatus) in flight, Plazas Islands, Galapagos
Flightless cormorant (Phalacrocorax harrisi) underwater, Punta Vicente Roca, Isabela Island, Galapagos, Ecuador, April
White-vented storm petrel (Oceanites gracilis galapagoensis) walking on surface of water whilst foraging, Galapagos
Marine iguana (Amblyrhynchus cristatus) on coast, Galapagos
Galapagos sea lions (Zalophus wollebaeki) fighting, Galapagos
Galapagos land iguana (Conolophus subcristatus) portrait with mouth open, Galapagos
Nazca booby (Sula granti) portrait, Galapagos, Ecuador
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Galapagos hawk (Buteo galapagoensis) landing, Galapagos, Ecuador. Vulnerable species
Galapagos hawk (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Galapagos hawk (Buteo galapagoensis) perched over sulphurous slope, Galapagos, Ecuador. Vulnerable species
People watching the Fissure eruption of Chico volcano into 9km diameter caldera. Sierra Negra volcano, Isabela Island, Galapagos, Ecuador. October 2005
Golden cownose rays (Rhinoptera steindachnyri) schooling in sheltered mangrove lagoon, Academy Bay, Santa Cruz Island, Galapagos, Ecuador
Yellow warbler (Setophaga petechia aureola) on lava rocks. Isabela Island, Galapagos. May
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Galapagos green turtle (Chelonia mydas agassizi) swimming in coastal waters of San Cristobal Island, Galapagos
Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Flightless cormorant (Phalacrocorax harrisi), adult and two chicks. Beagle Crater, Isabela Island, Galapagos
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Galapagos racer snakes (Pseudalsophis biserialis) group alert watching for prey on beach, Galapagos, June. Endangered species. Taken on location for BBC tv Blue Planet 2 series
Galapagos giant tortoise (Geochelone nigra porteri), Cerro El Chato, Santa Cruz Island, Galapagos Islands
Magnificent Frigate (Fregata magnificens) male displaying by inflating his throat pouch. North Seymour, Galapagos, Ecuador, April
Marine Iguana (Amblyrhynchus cristatus) basking on volcanic rock. Fernandina Island, Punta Espinosa, Galapagos, Ecuador, April
San Cristobal giant tortoises (Chelonoidis chatamensis), fighting, Galapaguera, San Cristobal Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) in the surf, Black Beach, Floreana Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) in water, Puerto Egas, Santiago Island, Galapagos
Galapagos lava lizard (Microlophus albemarlensis) sitting on Marine iguana (Amblyrhynchus cristatus) Punta Espinosa, Fernandina Island, Galapgos
Floreana giant tortoise hybrid descendants (Chelonoidis elephantopus) feeding on flower. These are descendants of the extinct Floreana giant tortoise, but are hybridised with other species
Galapagos green turtle (Chelonia mydas agassizii) swims over a coral reef. Darwin Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
