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Eastern rockhopper penguin (Eudyptes chrysocome filholi) mid-fall after jumping off rocky ledge to get to sea, surrounded by Bull kelp (Durvillaea antarctica), Auckland Island, Auckland Island
Ringed seal (Pusa hispida) resting on ice, Svalbard, Norway. April
Polar bear (Ursus maritimus) female with cub, walking over snow with mountains in background, Svalbard, Norway. April
Poppy bee (Osmia papaveris) resting on piece of Common poppy (Papaver rhoeas) petal, cut from the flower to line its nest
Red-footed tortoise (Chelonoidis carbonaria) portrait, Pantanal wetlands, Mato Grosso do Sul, Brazil
Chestnut-eared aracari (Pteroglossus castanotis) portrait, Pantanal wetlands, Mato Grosso, Brazil
Group of Little bee-eaters (Merops pusillus) perched side by side on branch in early morning, Allahein River, The Gambia
Two African scops owls (Otus senegalensis) perched in tree, Katavi National Park, Tanzania
Chamois (Rupicapra rupicapra) head portrait, Schwyz, Switzerland. November
Common toad (Bufo bufo) and Common frog (Rana temporaria) in attempted amplexus, Lucerne, Switzerland. April
Granular poison frog (Oophaga granulifera) resting on leaf at night, Uvita, Costa Rica
Jumping spider (Heliophanus sp. ) portrait, Lucerne, Switzerland. AprilJumping spider (Heliophanus sp.) portrait, Lucerne, Switzerland. April
Jumping spider (Hypositticus pubescens) male, resting on leaf, Lucerne, Switzerland. June. Focus stacked
Stoat (Mustela erminea) in winter coat, standing alert in grassland, Lucerne, Switzerland. March
Rainforest toad (Incilius aucoinae) male, portrait, Osa Peninsula, Costa Rica
Jumping spider (Lyssomanes sp. ) female, guarding its eggs on a leaf, Osa Peninsula, Costa Rica. Focus stackedJumping spider (Lyssomanes sp.) female, guarding its eggs on a leaf, Osa Peninsula, Costa Rica. Focus stacked
Two domestic Sheep (Ovis aries) standing side by side on moorland, Derbyshire, UK. August
Red deer (Cervus elaphus), young male, swimming through water. Bushy Park, London, UK. September
Waved albatross (Phoebastria irrorata) chick, aged two weeks, begging next to parent on nest. Punta Suarez, Espanola Island, Galapagos Islands, Ecuador
Short-eared owl (Asio flammeus) perched on branch. Galapagos Islands, Ecuador
Marine iguana (Amblyrhynchus cristatus) emerging from sea after feeding on algae. Fernandina Island, Galapagos Islands, Ecuador
Copan brook frog (Duellmanohyla soralia) resting on a leaf at night, Caribbean slope of Guatemala. Endangered
Group of California sea lions (Zalophus californianus) playing near the sea surface, Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
California sea lion (Zalophus californianus) close up with group playing in shallow water behind, Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Spectacled caiman (Caiman crocodilus) in shallow water, Amazon rainforest, Loreto, Peru
Wine throated hummingbird (Selasphorus ellioti) male, courtship display, Cloud forest, Atitlan Volcano, Solola, Guatemala
Reticulated tree frog (Dendropsophus reticulatus) portrait, Loreto, Peru
Little bittern (Ixobrychus minutus) perched in reedbed calling, Valkenhorst nature reserve, Valkenswaard, the Netherlands. May
Ruddy duck (Oxyura jamaicensis) male in courtship display. Valkenhorst nature reserve, Valkenswaard, The Netherlands. May
Long eared owl (Asio otus) perched on branch, Hampshire, UK. October. Captive, controlled conditions
Vervet (Chlorocebus pygerythrus) sitting on tree stump eating fruit, Uganda
Ring-tailed lemur (Lemur catta) head portrait. Captive, occurs in Madagascar, Endangered
Sand cat (Felis margarita) walking over sandy ground. Captive, occurs in North Africa, the Arabian Peninsula, Pakistan and the Middle East
Wild boar (Sus scrofa) piglet nuzzling a dog, La Pampa province, Patagonia, Argentina. Captive
Large hairy armadillo (Chaetophractus villosus) portrait, Peninsula Valdes, Patagonia, Argentina
Two banded plover (Charadrius falklandicus) female, incubating eggs on nest on pebbly beach, Cerro Avanzado protected area, Chubut, Patagonia, Argentina
Pichi / Dwarf armadillo (Zaedyus pichiy) in grassland, La Pampa Province, Patagonia, Argentina
Walrus (Odobenus rosmarus) male resting in water, Kapp Lee, Svalbard, Norway, Arctic Ocean, July
Female Olive ridley turtles (Lepidochelys olivacea) coming ashore to nest while others leaves, at sunrise, during massive arribada. Playa Escobilla Sanctuary, Oaxaca, Mexico. Pacific Ocean
Nilgiri tahr (Nilgiritragus hylocrius) standing next to road signage meant to warn vehicles about the possible road crossings of tahrs in the area, Valparai, Tamil Nadu, India
Lion-tailed macaque (Macaca silenus) infant, sitting among the troop with its finger in its mouth, Valparai, Tamil Nadu, India
Lesser whistling duck (Dendrocygna javanica) standing in long grass, Uran, Maharashtra, India
Small pearl-bordered fritillary butterfly {Boloria selene} head-on view, resting on grass, Cornwall, UK. June
Santa Catalina Island desert iguana (Dipsosaurus catalinensis) in front of Santa Catalina barrel cactus (Ferocactus diguetii), Santa Catalina Island, Loreto Bay National Park, Sea of Cortez, Mexico
Cape fur seal (Arctocephalus pusillus) juvenile resting on rock, Cape Cross Seal Reserve, Namibia
African desert elephant (Loxodonta africana), drinking from waterhole, Hoanib River, Damaraland, Namibia
Brown bear (Ursus arctos) walking across rock in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. September
Icelandic horse in flowering buttercup (Ranunculus sp. ) meadow, southwest Iceland. JuneIcelandic horse in flowering buttercup (Ranunculus sp.) meadow, southwest Iceland. June