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Close-up portrait of a Red Fox (Vulpes vulpes). Vosges, France, June
Chestnut arabian stallion running, California, USA
Pine marten (Martes martes) in tree trunk, Finland, July
Rough coated Jack Russell Terrier puppy, black, tan and white, portrait
Rough coated Jack Russell Terrier, tan and white, lying down, panting
Domestic dog, Old English Sheepdog / Bobtail, studio portrait
Pronghorn Antelope (Antilocapra americana) head portrait, South Dakota, USA
Grey Wolf (Canis lupus) head portrait of male, in snow, Captive
Portrait of a Bouvier des Flander cross-breed dog, sitting and panting
Cocker Spaniel head portrait, aged 5 years
Head portrait of a German Shepherd, panting
Golden Retriever puppy, 6 weeks, amongst gourds, Illinois, USA
Asiatic wild ass (Equus hemionus) with foal, captive, UK, endangered species
Orang utan baby (Pongo pygmaeus) head portrait, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
Orang utan (Pongo pygmaeus) baby hanging from trees, and chewing on fingers, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
Golden Retriever head portrait, lying in autumn leaves, panting
French Bulldog, portrait lying in autumn foliage
Burchells Zebra (Equus quagga burchellii) herd walking towards waterhole in the late afternoon, kicking up dust as they come. Etosha National Park, Namibia, Southern Africa
Large adult male Leopard (Panthera pardus) walking through the bush at night, Sabi Sand Private Game Reserve, South Africa, June
Black Jumping spider {Salticidae}. Tropical rainforest, Masoala Peninsula National Park, north east Madagascar
Snowy Owl (Bubo scandiaca) in flight over snow covered landscape, Ottawa, Ontario, Canada, December
A domestic pig (Sus scrofa domestica) swimming in the sea. Exuma Cays, Bahamas. Tropical West Atlantic Ocean. This family of pigs live on this beach in the Bahamas and enjoy swimming in the sea
Bobtail / Old English Sheepdog, sitting alongside a longhaired Chihuahua, tan and white coated
Mixed Breed dog running on grass, towards camera
Head portrait of Bornean Horned Frog (Megophrys nasuta) among the leaf litter in the lowland rainforest of Borneo. Danum Valley Conservation Area, Sabah, Malaysia
Stump-tailed Leaf Chameleon (Brookesia superciliaris) on white background, from rainforest leaf-litter. Masoala National Park, Madagascar
Polar bear (Ursus maritimus) sow with her cub outside their den in late winter, Arctic coast of Alaska
Red squirrel (Sciurus vulgaris) sitting upright in deep snow, Austria, Europe
Angler fish (Melanocetus murrayi) Mid-Atlantic Ridge, North Atlantic Ocean
Lionhead-cross rabbit sitting up on its haunches
Deepsea planktonic stage of crab development, Trondheimsfjord, North Atlantic Ocean
Pacific hatchetfish (Argyropelecus affinis) portrait
Giraffe (Giraffa camelopardalis) female bending down to calf, Okavango Delta, Botswana
Feral domestic rabbit (Oryctolagus cuniculus) juvenile running with dead leaf in mouth, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Raccoon (Procyon lotor) standing up investigating camera, portrait, Stanley park, Vancouver, British Columbia, Cananda, September
Asiatic lion cub (Panthera leo persica) looking up into tree, possibly at a bird, Gir Forest NP, Gujarat, India
Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills, Lake Kerkini, Greece, February. Highly Commended, 2011 Wildlife Photographer of the Year competition
Basset hound, portrait, resting head on back of car, eyes half closed, UK
RF- Grey seal (Halichoerus grypus) pups on Norfolk Beach, England, UK, December
Japanese macaque (Macaca fuscata fuscata) mother with rare white furred baby, Jigokudani Valley, Nagano Prefecture, Japan. June
RF- Juvenile Sumatran tiger (Panthera tigris sumatrae), aged four months, captive, occurs in Sumatra, Indonesia
Bengal Tiger (Panthera tigris) sub-adult, approximately 17-19 months old, amongst forest vegetation. Endangered. Bandhavgarh National Park, India. Non-ex
RF- Male Bengal Tiger (Panthera tigris tigris) face portrait in undergrowth, Kanha National Park, Madhya Pradesh, India. Endangered species
Saiga antelope (Saiga tatarica) male, standing in grassland in morning sunlight, Betpak Dala, Kazakhstan. Critically endangered
Galapagos giant tortoise (Chelonoidis nigra) portrait, Santa Cruz Island, Galapagos Islands, Ecuador. Endangered
Western grey kangaroo (Macropus fuliginosus) pair, male standing and female lying down, Kangaroo Island, Australia
Eastern rockhopper penguin (Eudyptes chrysocome filholi) mid-fall after jumping off rocky ledge to get to sea, surrounded by Bull kelp (Durvillaea antarctica), Auckland Island, Auckland Island
Ringed seal (Pusa hispida) resting on ice, Svalbard, Norway. April