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Bat-eared fox (Otocyon megalotis) walking, Namib-Naukluft National Park, Namib Desert, Namibia
Polar bear (Ursus maritimus) female coming out the den and shaking off snow. Wapusk National Park, Churchill, Manitoba, Canada, March
Hanuman langur / Northern Plains Grey Langur (Semnopithecus entellus) drinking from a water tap at Ranthambore fort. Ranthambore National Park, Rajasthan, India
Arctic Fox cub (Alopex / Vulpes lagopus) standing in fresh snow, blue morph. Dovrefjell National Park, Norway, September
Song thrush (Turdus philomelos clarkei) portrait, Hungary, May
A group of Pure Arab, Shagya Arab and East Bulgarian fillies running in snow, Kabiuk National Stud, Shumen, Bulgaria
Rough Collie, bitch, 9 months in straw
Japanese Macaques (Macaca fuscatata) grooming in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Black-backed Jackal (Canis mesomelas) drinking, Kgalagadi Transfrontier Park, South Africa. January
A portrait of a Mediterranean anthias (Anthias anthias) against open water. Secca Del Papa, Tavolara Marine Protected Area, Sardinia, Italy. Mediterranean Sea
A portrait of a nudibranch ( Limacia clavigera) searching for food on algae. Gulen, Norway. North East Atlantic Ocean
Portrait of a Spiny-chest Frog (Alsodes norae) endemic to Oncol Park, Chile, January
Yellow Labrador portrait in wild flowers
Spring lambs in meadow, UK, April
Gannet (Morus bassanus) adult portrait. Shetland Islands, Scotland, UK, September
RF- Hawk owl (Surnia ulula) in flight, Finland. March
RF- Red Squirrel (Sciurus vulgaris) in snow. Finland. February
RF- Gannet (Morus) in flight, Great Saltee, County Wexford, Ireland. April
RF- Black Grouse (Lyrurus tetrix) displaying. Finland. March
Coastal brown bear (Ursus arctos) fishing, Lake Clarke National Park, Alaska, September
Whooper swan (Cygnus cygnus) in flight in snowfall, Lake Kussharo, Japan, February
Coastal brown bear (Ursus arctos) resting, Lake Clarke National Park, Alaska, September
Marbled white butterfly (Melanargia galathea) nectaring on flower, Vallee de la Seille (Seille Valley), Lorraine, France, June
Kamchatka Brown Bear (Ursus arctos beringianus) stands on its back legs to more easily determine the source of an unfamiliar sound
Blue Maomao (Scorpis violaceus) shoal, Poor Knights Islands, New Zealand, February
Black rhino (Diceros bicornis) looking threatening, Nakuru National Park, Kenya
White rhino (Ceratotherium simum) in the rain, Nakuru National Park, Kenya
Great Grey owl (Strix nebulosa) taking off from the ground in snow, Finland, April. Bookplate from Danny Greens The Long Journey North
RF - Lemur Leaf Frog (Agalychnis lemur) in rainforest. Central Caribbean foothills, Costa Rica. IUCN Red List. Critically endangered species
Canary-shouldered thorn moth (Ennomos alniaria) portrait, Peak District National Park, Derbyshire, UK. October
Osprey (Pandion haliaetus) splashing water as it emerges from loch whilst fishing at dawn, Cairngorms National Park, Scotland, July
Portrait of Red Sea anemonefish (Amphiprion bicinctus) against open water. Nuweiba, Sinai, Egypt
RF- Northern Goshawk (Accipiter gentilis) male close-up, Scotland
A portrait of a small Candy stripe shrimp (Lebbeus grandimanus) in front of its host pink anemone (out of focus because of the magnification)
Ribbon sweetlips (Plectorhinchus polytaenia) USAT Liberty Wreck, Tulamben, Bali, Indonesia. Java Sea
Little egret (Egretta garzetta) shaking water off wings at sunset, Lake Csaj, Pusztaszer, Hungary. May. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Female Azure damselfly (Coenagrion puella), Vallee de la Seille (Seille Valley), Lorraine, France, June
Pine Grosbeak (Pinicola enucleator) male feeding on berries with traffic light behind, Oulu, Finland, December
Blue Tit (Paris caereleus) hanging below red berries on which it has been feeding, Uto Finland November
Reticulated Giraffe (Giraffa camelopardalis reticulata) portrait. Laikipia, Kenya
Southern elephant seal (Mirounga leonina) pup on beach, Caleta Valdes, Valdes Peninsula, Chubut, Patagonia, Argentina
Cheetah (Acinonyx jubatus) cub aged 6 months yawning, Masai-Mara Game Reserve, Kenya. Vulnerable species
RF- Head portrait of Giant panda (Ailuropoda melanoleuca) chewing on bamboo in snow, captive born in 2000. Occurs in China
RF- Mexican treefrog 1+Smilisca baudinii+2 on Spanish moss. Texas, USA
Pair of Common toads (Bufo bufo) in amplexus underwater, Belgium, March
Beluga whale (Delphinapterus leucas) swimming under ice and exhaling air, with scuba diver in background, Arctic circle Dive Center, White Sea, Karelia, northern Russia, captive
Atlantic Puffin (Fratercula arctica) resting on a snow bank, Isle of Hornoya, Varanger, Norway, April. Bookplate from Danny Greens The Long Journey North
White tailed Sea Eagle (Haliaeetus albicilla) in flight, Norway, July. Bookplate from Danny Greens The Long Journey North