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Capercaillie (Tetrao urogallus) in flight, Kuusamo, Finland, January
Portrait of a Yellow seahorse (Hippocampus guttulatus), Capo Galera, Alghero, Sardinia, Italy, Mediterranean Sea
Horsefly (Philipomyia aprica) Aosta Valley, Monte Rosa Massif, Pennine Alps, Italy. July
Zanzibar red colobus (Piliocolobus kirkii) in tree, Jozany Forest, Zanzibar, Tanzania. Highly endangered
White rhinoceros (Ceratotherium simum) mother and calf browsing. Entabeni Safari Conservancy, Limpopo region, Waterberg, South Africa, October
Snow Goose (Chen caerulescens), coming in to land. Bosque del Apache, New Mexico, USA, November
Polar bear (Ursus maritimus) adult walking over newly formed pack ice during fall freeze up, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska, USA
Long Horn cattle in meadow, Norfolk, UK July
A band of semi-feral Quarter mares and foals running across a creek, Estancia Don Amerigo, Chaco, Paraguay, January 2012
Close-up of Brown bear (Ursus arctos) in lake, Kamchatka, Far east Russia, August
Banded jawfish (Opistognathus macrognathus) male incubating eggs in mouth, which are at a late stage of development and will soon be ready for release
Yellow goatfish (Mulloidichthys martinicus) swimming above the reef, West End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Freerange Gloucester old spot domestic pig (Sus scrofa domestica) portrait with ears covering eyes, UK
Wild cat (Felis silvestris) in snowstorm captive, Germany
Hanuman Langur / Northern Plains Grey Langur (Semnopithecus entellus) mother with baby in tree. Kanha National Park, Madhya Pradesh, India
Mute swan (Cygnus olor) on water, Wheatfen, Norfolk, UK, November
Jumping spider (Euophrys frontalis) male amongst flower petals, Peak District National Park, Derbyshire, UK, June
Harvest mouse (Micromys minutus soricinus) feeding on blackberry, captive, Dorset, UK, October
Roe deer (Capreolus capreolus) at sunset, Vosges, France, August
Tropical Butterfly (Adepha) on a leaf with wings spread. Danum Valley, Sabah, Borneo, Malaysia
Portrait of a Lynx (Lynx lynx) mother and kitten. Hanau, Germany, July. Captive
Common green tree frog (Litoria caerulea) in rainforest, Iron Range National Park, Cape York Peninsula, North Queensland, Australia, October
Indian / Asian one-horned rhinoceros (Rhinoceros unicornis) approaching, Kaziranga National Park, Assam, India
Grey seal (Halichoerus grypus) adult female among breaking waves, Lincolnshire, UK Non-ex
Young windmill-eared rabbit and matching guinea-pig
Blue Bearded Collie puppy, 3 months, yawning
Ginger kitten, 7 weeks, sitting between two young Lionhead-Lop rabbits
Border Collie bitch, running towards the camera
Black Rhinoceros (Diceros bicornis) charging. Etosha National Park, Namibia, January
West Highland White Terrier sitting
Playful Border Collie puppies, 6 weeks
Border Collie puppy portrait
Domestic dog, Border Collie lying on grass, panting
Jumping Spider (Salticidae) perched on Bigtooth Maple leaf (Acer grandidentatum). Lost Maples State Park, Hill Country, Central Texas, USA, November
Close-up portrait of a Red Fox (Vulpes vulpes). Vosges, France, June
Konik horses (Equus caballus) - wild Konik filly, Millingerwaard nature reserve, Netherlands, April
White-lipped / Green tree viper (Cryptelytrops albolabris) captive, from SE Asia
Portrait of a Common Racoon (Procyon lotor). Stanley Park, Vancouver, Canada, March
Brown bear (Ursus arctos) strolling along the beach in the rain, looking towards the camera. Lake Clark National Park, Alaska, USA, July
Portrait of an Emerald Glass Frog (Espadarana / Centrolenella prosoblepon). Captive. Choco Region of northwest Ecuador
Southern plains grey / Hanuman langur {Semnopithecus dussumieri} two adolescents sitting, Thar desert, Rajasthan, India
Red Deer (Cervus elaphus) male standing in snow. The Netherlands, January
Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January
Portrait of Grey Heron (Ardea cinerea) looking directly at lens. The Netherlands, June
Brown Hare (Lepus europaeus) portrait gazing alertly. Perthshire, Scotland, June
Domestic dog, Scottish Terrier / Aberdeen Terrier, studio portrait
Brown bear (Ursus arctos) running through water, fishing, Kamchatka, Far east Russia, August
Grey Wolf (Canis lupus) head portrait of male, sleeping in snow, Captive