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Brown Bear (Ursus arctos) mother and 3-4 month cubs foraging for clams. Katmai National Park, Alaska, USA, June
White ibis (Eudocimus albus) flock in breeding plumage, backlit against dark background, four individuals flying in to join flock, Tampa Bay, Florida, USA, March
Four Yellow-billed storks (Mycteria ibis) standing in water, Lake Nakuru, Nakuru National Park, Kenya
Herd of Impala (Aepyceros melampus) silhouetted at sunset, Ngorongoro Conservation Area, Tanzania
Atlantic puffins (Fratercula arctica) at dusk, Fair Isle, Shetland, July. Bookplate from Danny Greens The Long Journey North
Sequence of a breaching Southern right whale (Eubalaena australis) Golfo Nuevo, Peninsula Valdes, UNESCO Natural World Heritage Site, Chubut, Patagonia, Argentina, Atlantic Ocean
Whooper Swan (Cygnus cygnus) on water with cygnets. Iceland, June
Wolves (Canis lupus) sitting on a hillside in snow. Yellowstone, USA, February
Lion (Panthera leo) males resting, Masai-Mara Game Reserve, Kenya. Vulnerable species
Great skua (Stercocarius skua) in flight over Gannet (Morus bassanus) colony, looking for small chicks, whilst Gannets call defensively, Shetland Isles, July
Dalmatian Pelicans (Pelecanus crispus) portrait. Lake Kerkini, Greece, March 2012
Cheetah (Acinonyx jubatus) cubs aged 6 / 7 weeks old playing, Masai-Mara Game Reserve, KenyaCheetah (Acinonyx jubatus) cubs aged 6/7 weeks old playing, Masai-Mara Game Reserve, Kenya. Vulnerable species
Honeybee workers exchanging food - known as trophallaxis (Apis mellifera) Sussex, UK
Hand reared Whooper Swans (Cygnus cygnus) in flight. Birds imprinted and trained by Lloyd and Rose Buck for filming purposes. Somerset, UK, January 2011
Killer whale / Orca (Orcinus orca) surfacing with three seabirds flying, Tysfjord, Norway, November
Small flock of Greater flamingo (Phoenicopterus roseus) wading on inland Kalloni lake, Lesbos, Greece
Four Crowned cranes {Balearica regulorum} Kenya
Four Yellow-eyed penguins (Megadyptes antipodes) walking up a sand dune towards their nests, Otago Peninsula, Otago, South Island, New Zealand. September. Endangered Species
Four Galapagos sea lions (Zalophus wollebaeki) sleeping on beach, Floreana Island, Galapagos, Endangered species