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Autumnal Quaking aspen tree (Populus tremuloides) at Nares Lake, with Montana Mountain beyond, near Carcross, Yukon Territories, Canada, September 2013
Lake Baikal landscape in autumn, trees reflected in water. Zabaikalsky National Park, Siberia, Russia, September 2013
Autumnal Beech (Fagus) trees, Savernake Forest, Wiltshire, UK, November 2012
Red squirrel (Sciurus vulgaris) foraging in forest, climbing down tree trunk, Cairngorms National Park, Scotland, October
Wild cat (Felis silvestris) climbing tree in autumn, Black Forest, Baden-Wurttemberg, Germany. October
Landscape of the Verzasca River in autumn, Canton Tessin, Switzerland, November 2011. Taken for the Freshwater Project
Toadstoosl on tree stump in forest, Vosges, France, November 2013
Misty Beech (Fagus sylvatica) forest in autumn, Vosges mountains, France, November 2013
Sheathed Woodtuft fungi (Kuehneromyces mutabilis) Surrey, England, UK, November
Partially wooded uplands with Five Sisters of Kintail in background, Northwest Highlands, Scotland, UK.September
Collared earth star (Geastrum triplex), two on woodland floor. South Yorkshire, England, UK. September
Eurasian otter (Lutra lutra) feeding whilst on log in woodland burn, Glenfeshie, Cairngorms National Park, Scotland, UK, September
Weasel (Mustela nivalis) investigating birch stump with bracket fungus in autumn woodland, Sheffield
Red Squirrel (Sciurus vulgaris), at woodland pool, Cairngorms National Park, Scotland, UK.November
Stunted oak woodland covered in moss, Wistmans Wood, Devon, UK. August. Highly Commended in the Wild Woods category of the British Wildlife Photography Awards (BWPA) competition 2013
Andean Emerald Hummingbird (Amazilia franciae), Mindo cloud forest area, Ecuador, July
Red shanked douc langur (Pygathrix nemaeus) Son Tra Nature Reserve, Vietnam
Black woodpecker (Dryocopus martius) male perched on tree trunk, trees covered in hoar frost. Kuusamo, Northern Ostrobothnia, Finland. February
Stream in spate in native oak woodland in summer, Clonaig, Kintyre, Argyll, Scotland, UK, July 2015
Montane rainforest, Hienghene, New Caledonia
Rainforest and river outside the Batak village of Sitio Manggapin in Cleopatra sRainforest and river outside the Batak village of Sitio Manggapin in Cleopatras Needle Critical Habitat, Palawan, Philippines
Coral formations at low tide in Prony Bay in the Southern Lagoon, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Temmincks tragopan (Tragopan temminckii) Tangjiahe Nature Reserve, Sichuan province, China
Tapanuli orangutan (Pongo tapanuliensis) Togus, adult flanged male, trying to nap in tree, Batang Toru Forest. Sumatran Orangutan Conservation Project, North Sumatran Province, Indonesia
Puente de Dios cascades, Tamasopo, Huasteca Region, Mexico, August
Herald moth (Scoliopteryx libatrix) camouflaged in leaf litter, Gosford Forest Park, Co. Armagh, Northern Ireland. September
Mountain gorilla (Gorilla beringei beringei), dominant silverback in forest clearing. Volcanoes National Park, Rwanda
Beech woodland near Blackwater Brook. New Forest National Park. November 2014
Tapanuli Orangutan (Pongo tapanuliensis) Beti, juvenile female approximate age 6 years, playing with mother, Beta, Batang Toru Forest, Sumatran Orangutan Conservation Project
Ivy (Hedera helix) covered tree trunks in Knight Wood, Hampshire, UK, October
Synchronous fire flies (Pteroptyx sp. ) mangroves along the Kinabatangan River, Sabah, BorneoSynchronous fire flies (Pteroptyx sp.) mangroves along the Kinabatangan River, Sabah, Borneo
Montane horned frog (Megophrys kobayashii) in the leaf litter at night. Mount Kinabalu (around 1400m asl), Sabah, Borneo
Giant honey bee nest (Apis dorsata) up in a giant Mengaris tree (Koompassia excelsa) dipterocarp forest in Danum Valley, Sabah, Borneo, Malaysia
Spirit / Kermode bear (Ursus americanus kermodei) catching fish in river, most likely salmon, the Great Bear Rainforest, British Columbia, Canada, October
Kaieteur Falls is the worlds widest single drop waterfall, located on the Potaro river in the Kaieteur National Park, in Essequibo, Guyana, South America
Grass snake (Natrix natrix) in habitat, Aland Islands, Finland, May
Roe deer (Capreolus capreolus) young male walking through snow. Close-up, portrait. Norway, February
Suspension bridge across the river, entrance to Gunung Mulu National Park UNESCO Natural World Heritage Site, Malaysian Borneo
Willow (Salix) plantation, Cuenca, Castilla-La Mancha, Spain, December
Bornean orangutan (Pongo pygmaeus) male, Gunung Palung National Park, Borneo
Leaf mimic bush cricket or katydid (unknown species, family Tettigoniidae) in the middle of moulting its skin. Moulting from one sub-adult stage (instar) to another is known as ecdysis
White fronted capuchin monkey (Cebus capucinus) feeding, March, Costa Rica
River through Virgin Komi Forests UNESCO World Heritage site, the largest virgin forests in Europe. Ural Mountains, Komi Republic, Russia. August 2016
Eastern Ring-tailed Vontsira (Galidia elegans elegans) (endemic family Eupleridae) foraging on forest floor. Marojejy National Park, north east Madagascar
Female Red-eyed Tree Frog (Agalychnis callidryas) - Caribbean slope race (blue flanks) - obscured in rainforest foliage. Mid-altitude rainforest near Aranal, central Costa Rica
Pristine tropical rainforest and granite cliffs in mist, Morne Seychelles National Park, Mahe Island, Republic of Seychelles
Brazilian tapir (Tapirus terrestris) portrait, Tambopata Research Centre, Peru
Guianan cock-of-the-rock (Rupicola rupicola), two males fighting during lek, Guianan Shield, Suriname