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Kinabalu giant red leech (Mimobdella buettikoferi) feeding on Kinabalu giant earthworm (Pheretima darnleiensis), on Mount Kinabalu, Borneo
Proboscis monkey (Nasalis larvatus) female feeding, Kinabatangan River, Sabah, Borneo
Mountain tree shrew (Tupaia montana) feeding on nectar secreted by the endemic Pitcher Plant (Nepenthes kinabaluensis) Montane forests (at 2200m-3000m), slopes of Mt Kinabalu
Sumatran pit viper (Trimeresurus sumatranus) female in rain forest understorey. Danum Valley, Sabah, Borneo
Red deer (Cervus elaphus) stag in autumn woodlands, Bradgate Park, Leicestershire, UK, October
RF - Red Deer stags (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
RF - European Badger (Meles meles) foraging at tree trunk in deciduous woodland. Mid Devon, UK. June
Red deer (Cervus elaphus) stag in the snow, Scotland, March
Pair of Sockeye salmon (Oncorhynchus nerka) on their redd in a shallow stream. Female in front of male. Huihil Creek, Adams River, British Colombia, Canada, October
A climber plant in the montane rainforest, near FakFak, Mainland New Guinea, Western Papua, Indonesian New Guinea
Emerald Glass Frog (Centrolenella proseblepan) on leaf. Mid-altitude rainforest, Bosque de Paz, Pacific slope, Costa Rica, Central America
Forest reindeer, (Rangifer tarandus fennicus) in velvet, Viiksimo, Kuhmo region. Finland, July. Rare subspecies which were nearly extinct in the 19th Century
River Plym flowing fast through Dewerstone Wood, Shaugh Prior, Dartmoor National Park, Devon, England, UK. May 2015
Beech (Fagus sylvatica) forest in Tarcu mountains nature reserve, Natura 2000 area, Southern Carpathians, Romania, May 2014
African forest elephant (Loxodonta cyclotis) in water, Lekoli River, Republic of Congo (Congo-Brazzaville), Africa. Vulnerable species
Eurasian lynx (Lynx lynx) climbing tree, Black Forest, Baden-Wurttemberg, Germany. November. April. Captive
Fer-de-lance (Bothrops asper) camouflaged on the rainforest floor, Corcovado National Park, Osa Peninsula, Costa Rica
Satanic leaf-tailed gecko (Uroplatus phantasticus) on twig, Andasibe-Mantadia National Park, Alaotra-Mangoro Region, Madagascar
Large aerial pitchers of Veitchs pitcher plant (Nepenthes veitchii) growing up a tree trunk. Maliau Basin, Borneo
Melanistic leopard / Black panther (Panthera pardus) on territorial patrol on track, Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India
Silverback Mountain gorilla (Gorilla beringei) in the bamboo forest, this is Munyinya the leader of Hirwa group, Sabyinyo volcano, Volcano National Park, Rwanda
Bamboo forest, Shikoku Mura open air museum, Takamatsu, Shikoku Island, Japan, May
Badger (Meles meles) foraging in woodland on edge of forest, Black Forest, The Black Forest, Germany, May. Highly commended in Mammals category, GDT competition 2011
Beech woodland in dense fog. Derbyshire, UK, November
Palm trees (Licuala valida) in swamp forest, Lambir NP, Borneo, Sarawak, Malaysia
Moose (Alces alces) head portrait of bull standing in forest clearing, Cap Breton Highlands National Park, Nova Scotia, Canada, September
Wood Anemones (Anemone nemorosa) carpeting a woodland floor, low angle fish eye view, Peak District National Park, Derbyshire, UK, May
Jaguar (Panthera onca) portrait, Tambopata, Madre de Dios, Peru
Ecuadorian toad (Rhaebo ecuadorensis) on branch, Yasuni National Park, Orellana, Ecuador
Sumatran orangutan (Pongo abelii) female with very young baby, Gunung Leuser National Park, Sumatra, Indonesia
Red eyed tree frog (Agalychnis callidryas) La Selva Field Station, Costa Rica
The San Rafael waterfall, the biggest falls in Ecuador, located on the boundary of the Amazon with The Andes. San Rafael, Napo, Ecuador. February 2016
Strawberry poison dart frog (Oophaga pumilio) La Selva Field Station, Costa Rica
Strawberry poison dart frog (Oophaga / Dendrobates pumilio) sitting in cup fungus, La Selva Field Station, Costa Rica
Forest with mist on the slopes of Mount Lidgbird. Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia
Parsons Chameleon (Calumma parsonii) climbing in rainforest understorey. Andasibe-Mantadia National Park, eastern Madagascar
Las Golondrinas Waterfalls, Montes Azules Biosphere Reserve, Chiapas. Mexico, March 2017
Misol-ha Waterfall, Ejido San Miguel, Salto de Agua Municipality, Chiapas. Mexico, March 2017
Long-tailed macaque (Macaca fascicularis) in a fruiting strangler fig tree (Ficus dubia). Gunung Palung National Park, Borneo
Hercules moth (Coscinocera hercules) recently emerged in montane rinforest. Ambua Lodge, Tari, Hela Province, Papua New Guinea. June
Oxbow lake adjacent to the Karawari River, Papua New Guinea, August
Black-crowned Central American squirrel monkey (Saimiri oerstedii oerstedii) Osa Peninsula, Costa Rica. IUCN Red List Vulnerable species
Scarlet macaw (Ara macao) pair in tree, Osa Peninsula, Costa Rica
RF - Scarlet macaw (Ara macao) pair preening, Osa Peninsula, Costa Rica
RF - Eyelash Pit Viper (Bothriechis schlegelii) Costa Rica
Tualang / Mengaris tree (Koompassia excelsa) emerging from canopy amongst cloud. Danum Valley, Sabah, Borneo, May 2011
Cup fungi (Cookeina sp) growing on decaying log, Danum Valley, Sabah, Borneo
