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Adult bull African Elephant (Loxononta africana) reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October
RF- Contorted trunks of Beech trees (Fagus sylvatica) in autumn mist. Beacon Hill Country Park, National Forest, Leicestershire, UK. October 2010
Red squirrel (Sciurus vulgaris) climbing tree trunk with autumn leaves, Highlands, Scotland, October 2015
Red squirrel (Sciurus vulgaris) on branch in autumnal forest, Cairngorms National Park, Highlands, Scotland, UK, October
Red squirrel (Sciurus vulgaris) silhouetted against autumnal woodland, Highlands, Scotland, UK, October
Red squirrel (Sciurus vulgaris) on forest floor with autumn leaves Highlands, Scotland, UK, October
Silver birch (Betula pendula) woodland in early autumn, Craigellachie National Nature Reserve, Aviemore, Cairngorms National Park, Scotland, UK, October 2014
Tropical Screech Owl (Megascops choliba) perched in tree hole in rainforest canopy. Cristalino State Park, Amazonia, Brazil, South America
European robin (Erithacus rubecula) adult with autumn leaves in background, Punkaharju, Etela-Savo / Southern Savonia, Ita-Suomi / Eastern Finland, Finland. September
Chestnut-headed Tesia (Tesia castaneocoronata) perched, Sichuan Province, China
Crested drongo (Dicrurus forficatus) rear view, Berenty Reserve, Madagascar
Maple leaves backlit on lightbox, Broxwater, Cornwall, UK. November 2013
Autumnal foliage of Fireweed (Epilobium angustifolium) Kluane National Park, Yukon Territories, Canada, September
Okavango robbers (Rhabdalestes maunensis) fish swimming up the temporary Selinda Spillway, northern Botswana. This shallow river cuts across the Kalahari Desert
Geranium procurrens leaf, Norfolk garden, UK, June
Malayan tiger (Panthera tigris jacksoni), Malaysia. Captive. An Endangered species, only around 500 remain
Crested Owl (Lophostrix cristata) lowland rainforests near Cristalino Jungle Lodge, Cristalino State Park, Brazil
Harts tongue fern (Phyllitis scolopendrium) leaf unfurling, Cornwall, UK, May
Shining honeycreeper (Cyanerpes lucidus) Costa Rica
Morel fungus (Mitrophora semilibera) among leaf litter. Lorraine, France, March
Iberian black pig foraging in oak woodland, Sierra de Aracena Natural Park, Huelva, Andalucia, Spain, Europe. Breed used to produce Iberico ham / Jamon Iberico
Vanzolinis Spiny-chest Frog 1+Alsodes vanzolinii+2 on Eucalyptus leaves, endemic to the Nahuelbuta mountain range, Chile, December. Critically Endangered species
Common Cuscus (Phalanger orientalis) captive occurs in Timor to New Guinea. Captive
Rhinoceros beetle (Chalcosoma moellenkampi) climbing up tree trunk, Danum Valley Reserve, Sabah, Borneo, Malaysia
Phayers Leaf monkey (Trachypithecus phayrei) Gaoligongshan NP, Yunnan province, China
Male Panther Chameleons (Furcifer pardalis) fighting. Nosy Be, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, January 2010
Aquatic Warbler (Acrocephalus paludicola). Zvanets fen mire, Belarus, Europe. Vulnerable Species
RF- Backlit Fern (Philotum) growing among bluebells (Endymion nonscriptus). Lanhydrock, Cornwall, UK. May
RF- Beech (Fagus) and Bluebell (Hyacinthoides non-scripta) woodland at Lanhydrock, Cornwall. UK
Canal Zone tree frog (Hypsiboas rufitelus) La Selva, Costa Rica
Female Jaguar (Panthera onca), captive, occurs in Southern and Central America
Mist in the cloud forest at Sombom Ridge, Saruwaged Range, Huon Peninsula, Papua New Guinea
European wild cat (Felis silvestris) kitten portrait, Bavarian Forest National Park, Germany, captive
Marbled white butterfly (Melanargia galathea) nectaring on flower, Vallee de la Seille (Seille Valley), Lorraine, France, June
Ornate rainfrog (Pristimantis ornatissimus) on leaf, Otongachi, Santo Domingo de los Tsachilas, Ecuador, Vulnerable species
Coastal glassfrog (Cochranella litoralis) on leaf with eggs. San Lorenzo, Esmeraldas, Ecuador
RF- Sunrise and mist over lowland dipterocarp rainforest. Danum valley, Sabah, Borneo, Malaysia
Lemur leaf frog (Agalychnis lemur) Central Caribbean foothills, Costa Rica. IUCN Red List critically endangered species
RF - Lemur Leaf Frog (Agalychnis lemur) in rainforest. Central Caribbean foothills, Costa Rica. IUCN Red List. Critically endangered species
Black headed gull (Chroicocephalus ridibundus) taking flight, artistically blurred photograph. Cheshire, UK, October. Highly honoured in the Art in Nature category of the Natures Best Windland Smith
RF - Small pearl-bordered fritillary (Boloria selene) butterfly, Marsland mouth, North Devon, UK. May
Reflections of tree in forest floor after rain, Blue River Provincial Park / Parc Provincial de la Riviere Bleue, New Caledonia
Weasel (Mustela nivalis) head and neck looking out of yellow autumn acer leaves, Sheffield, England, UK
Red-eyed Treefrog (Agalychnis callidryas) on leaf, captive, from South America
RF - Black panther / melanistic Leopard (Panthera pardus) peering through leaves, captive
Ash (Fraxinus excelsior) fresh leaf in spring, Dorset, UK April
Norfolk Terrier portrait sitting in autumn foliage
