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A female Spiny Seahorse (Hippocampus guttulatus) in weeds. Sardina, Gran Canaria, Canary Islands, East Atlantic Ocean, October
A male Weedy Seadragon (Phyllopteryx taeniolatus) carrying eggs along tail swims through a Kelp (Macrocystis pyrifera) forest in Tasmania, Australia
A bull California sealion (Zalophus californianus) bursting through a school of baitfish in the shallows. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean
A long exposure of a Tiger Shark (Galeocerdo cuvier) portrait, at the surface at dusk. Little Bahama Bank, Bahamas. West Atlantic Ocean. March
Manta ray (Manta birostris) feeding on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean
Lemon shark (Negaprion brevirostris) at the surface, split level. Little Bahama Bank, Bahamas. West Atlantic Ocean
In-camera motion blur image of a female Yellow Seahorse (Hippocampus kuda). Lembeh Strait, Sulawesi, Indonesia, Molluca Sea, March
A batch of Leafy Seadragon (Phycodurus eques) eggs on the tail of a male. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
Female Slingjaw wrasse {Epibulus insidiator} amongst coral, Red Sea, Egypt
Large / Pot Bellied Seahorse (Hippocampus abdominalis) face portrait. Blairgowrie Marina, Mornington Peninsular, Victoria, Australia, March
Two Golden / Masked butterflyfish, Red Sea, Eygpt
Close up of the scale details of a Yellow-mask angelfish (Pomacanthus xanthometopon). Baa Atoll, Maldives. Indian Ocean
Lyretail anthias / Goldies (Pseudanthias squamipinnis) over coral reef with soft corals (Dendronephthya sp) Egypt, Red Sea
Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October
RF - Titan triggerfish (Balistoides viridescens) swimming over hard coral (Acropora sp.) gardens. Kri Island, Raja Ampat, West Papua, Indonesia. Dampier Strait, tropical west Pacific Ocean
Detail of pattern of an adult Emperor angelfish (Pomacanthus imperator) Seraya, Tulamben, Bali, Indonesia. Java Sea
Blotcheye / Bigscale Soldierfish (Myripristis berndti), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Pygmy seahorse, not currently described hiding in sea fan, Raja Ampat, Irian Jaya, West Papua, Indonesia, Pacific Ocean
Rock Beauty (Holacanthus tricolor). Cancun National Park, Caribbean Sea, Mexico
Gorgonian seahorse (Hippocampus bargibanti) camouflaged in Gorgonian, Lembeh Strait, North Sulawesi, Indonesia
Close-up of a Common crocodilefish / Carpet flathead (Papilloculiceps longiceps) camouflaged on sea bed, coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea
Close up of head of European Plaice (Pleuronectes platessa) camouflaged in sand.Taken in mobile field studio. Isle of Mull, Scotland, UK. June
Outline of an Angelshark (Squatina squatina) buried in the sand. Gran Canaria, Canary Islands, Spain. East Atlantic Ocean
A pair of Leafy Seadragons (Phycodurus eques) camouflaged sheltering amongst seaweeds. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, December
Giant frogfish {Antennarius commersoni} on Mushroom Leather coral, Sulawesi, Indonesia
Schoolmaster snapper (Lutjanus apodus) school in the reef, Little Cayman island, Cayman Islands, Caribbean
Black grouper (Mycteroperca bonaci) and Caribbean Reef Shark (Carcharhinus perezi), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba, January
Bridled parrotfish (Scarus frenatus) clownish grin reveals its power tools: grinding teeth used to scrape algae from rock, Maldives, Indian Ocean
Split image of Blue shark (Prionace glauca) close to the surface, Pico Island, Azores, Portugal, Atlantic Ocean, July
Shoal of Longfin bannerfish (Heniochus acuminatus) Maldives, Indian Ocean
Shoaling glassfish (Parapriacanthus sp. ) shoal, Maldives, Indian OceanShoaling glassfish (Parapriacanthus sp.) shoal, Maldives, Indian Ocean
Portrait of a Yellow seahorse (Hippocampus guttulatus), Capo Galera, Alghero, Sardinia, Italy, Mediterranean Sea
Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
A school of lesser devil rays (Mobula hypostoma) flying through sunbeams as they feed. Isla Mujeres, Quintana Roo, Yucatan Peninsula, Mexico. Caribbean Sea
Whitetip reef sharks (Triaenodon obesus) pack silhouetted following scent trail in water column, Cocos Island, Costa Rica, Pacific Ocean
Emperor angelfish (Pomacanthus imperator) with soft corals and sweepers. Egypt, Red Sea
Little egret (Egretta garzetta) attempting to catch fish, Camargue, France, July
Brown trout (Salmo trutta) resting on river bed near the Aigas Field Studies Centre, Inverness-shire, Scotland, July
Spotted scorpionfish (Scorpaena plumieri) lying motionless and camouflaged in soft corals. West Palm Beach, Gulf Stream, West Atlantic Ocean, Florida, USA
Banded jawfish (Opistognathus macrognathus) male incubating eggs in mouth, which are at a late stage of development and will soon be ready for release
Yellow goatfish (Mulloidichthys martinicus) swimming above the reef, West End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Grey Heron (Ardea cinerea) taking a fish from water. Kiskunsagi National Park, Hungary, May
Manta Ray (Manta Birostris) and Devil Ray (Mobula tarapacana) in background, accompanied by Pilot Fish (Naucrates ductor) and Remora. Santa Maria Island, Azores, September
Blue Shark (Prionace glauca) at the sea surface. Santa Maria, Azores, August
Blue Sharks (Prionace glauca) at the sea surface. Santa Maria, Azores, August
Caribbean reef shark (Carcharhinus perezi) Jardines de la Reina National Park, Cuba
Great Blue Shark (Prionace glauca), dorsal view against dark water. Over 20 million Blue Sharks are fished annually, as bycatch and due to the high value of their fins
A portrait of a Leafy Seadragon (Phycodurus eques). Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
