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Red / Blotcheye soldierfish (Myripristis murdjan) and sea whips coral, Junceella at Fotteyo caves, Maldives, Indian Ocean
Red Sea anemonefish (Amphiprion bicinctus) in sea-anemone, Red Sea
Long-beaked Coralfish (Chelmon rostratus) eye, Inanuran Island, Danajon Bank, Central Visayas, Philippines, April
Rainbow trout (Oncorhynchus mykiss) under trees, Capernwray Lake, Lake District, England, UK
Blue Maomao (Scorpis violaceus) shoal, Poor Knights Islands, New Zealand, February
Sargassum fish (Histrio histrio) swimming with discarded fishing net, Dominica
Osprey (Pandion haliaetus) with fish prey on feeding perch in the rain, Cairngorms National Park, Scotland, UK, July
Kingfisher (Alcedo atthis) flying up from water with caught fish, Worcestershire, UK, March
Golden damselfish (Amblyglyphidodon aureus) hiding amongst the branches of red whip coral (Ctenocella sp.) Misool, Raja Ampat, West Papua, Indonesia
Pair of Giant frogfish (Antennarius commersoni), male on the left, female above on right on sponges. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Tiny (10mm) pygmy seahorse (Hippocampus bargibanti) sheltering in seafan (Muricella sp)Tiny (10mm) pygmy seahorse (Hippocampus bargibanti) sheltering in seafan (Muricella sp.) Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Translucent goby (Bryaninops erythrops) rests on sponge (Pione vastifica) covered in spionid worms (Polydorella smurovi) Nuweiba, Sinai, Egypt. Gulf of Aqaba, Red Sea
Portrait of Red Sea anemonefish (Amphiprion bicinctus) against open water. Nuweiba, Sinai, Egypt
Female Lionfish (Pterois volitans) on coral reef. Jackfish Alley, Ras Mohammed Marine Park, Sinai, Egypt. Gulf of Aqaba, Red Sea
Close up detail photo of a Spotted moray (Gymnothorax isingteena), Baa Atoll, Maldives. Indian Ocean
Cleaner wrasse (Labroides dimidiatus) swims out from the gill of a Blue spotted pufferfish (Arothron caeruleopunctatus). East of Eden, Similan Islands, Thailand. Andaman Sea, Indian Ocean
Longfin batfish (Platax teira) with Fusiliers (Caesio spp.) in the current in front of Baby Rock pinnacle. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea
Clarks anemonefish (Amphiprion clarkii) portrait in its host Bubble-tip anemone: Entacmaea quadricolor). Horseshoe Bay, Rinca Island, Komodo National Park, Indonesia
Vibrant Red Sea reef scene, with orange female Scalefin anthias (Pseudanthias squamipinnis) swarming in front of Fire coral (Millepora dichotoma) feeding on plankton brought to the reef by currents
A long exposure renders Bar jacks (Carangoides ruber) as ghostly trails as they swim around a sponge, on a coral reef, at night. East End, Grand Cayman, Cayman Islands
Eye detail of Topknot (Zeugopterus punctatus) flatfish, Loch Fyne, Argyll, Scotland, UK, July
Portrait of pair of Bohar snappers (Lutjanus bohar) in their spawning aggregation. Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Portrait of male Scalefin anthias (Pseudanthias squamipinnis) swimming in open water above coral reef, with bokeh effect in water from the sunset. The Alternatives, Sinai, Egypt
Indigo hamlets (Hypoplectrus indigo) pair spawning at dusk, George Town, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Schoolmaster snapper (Lutjanus apodus) school, under a coral ledge, Snapper Hole, East End, Grand Cayman, Cayman Islands, Caribbean Sea
A pair of captive Common / Short snouted seahorses (Hippocampus hippocampus) in aquarium, Weymouth Sealife Centre, Dorset, England, UK
Group of Circular spadefish (Platax orbicularis) gather at a cleaning station beneath a table coral (Acropora sp.) on a coral reef. Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea
RF - Roving coralgrouper (Plectropomus pessuliferus) is cleaned by Bluestreak cleaner wrasse (Labroides dimidiatus). Ras Mohammed Marine Park; Sinai, Egypt. Red Sea
Green jacks (Caranx caballus) long exposure, La Paz, Mexico. Gulf of California, Sea of Cortez, East Pacific Ocean
Red gorgonians (Lophogorgia chilensis) growing beneath a Giant kelp (Macrocystis pyrifera) forest, with female California sheephead (Semicossyphus pulcher). Santa Barbara Island, Channel Islands
RF - Shallow depth of field portrait of male Scalefin anthia (Pseudanthias squamipinnis) swimming in open water above coral reef photographed against sunset. The Alternatives, Sinai, Egypt
RF - Scissortail sergeants fish (Abudefduf sexfasciatus) sheltering beneath hard coral (Acropora sp)RF - Scissortail sergeants fish (Abudefduf sexfasciatus) sheltering beneath hard coral (Acropora sp.) in the evening on a coral reef, Red Sea
Caribbean reef sharks (Carcharhinus perezi) swimming through the middle of a school of yellowtail snappers (Ocyurus chrysurus). Grand Bahama, Bahamas. Tropical West Atlantic Ocean
Colourful reef scene with Green tubastrea (Tubastrea micrantha), Soft corals (Scleronephthya sp)Colourful reef scene with Green tubastrea (Tubastrea micrantha), Soft corals (Scleronephthya sp.) and Scalefin anthias (Pseudanthias squamipinnis) and Magenta slender anthias (Luzonichthys waitei)
Redtail butterflyfish (Chaetodon collare) pair in front of a smooth Sea fan (Annella mollis). Butterflyfish regularly form stable pairs. Similan Islands, Thailand. Andaman Sea, Indian Ocean
Coral grouper (Cephalopholis miniata) guarding its territory on a colourful coral reef. East Of Eden, Similan Islands, Thailand. Andaman Sea, Indian Ocean
Spotlight goby (Gobiosoma louisae) looking out from inside a Yellow tube sponge (Aplysina fistularis), growing on a coral reef. Bloody Bay Wall, Little Cayman, Cayman Islands, British West Indies
Long exposure of a juvenile Harlequin sweetlips (Plectorhinchus chaetodonoides), which dance as a defence from predators. The dance may mimic the movements of poisonous flatworms or may simply
Ribbon sweetlips (Plectorhinchus polytaenia) USAT Liberty Wreck, Tulamben, Bali, Indonesia. Java Sea
RF - Bluestreak cleaner wrasse (Labroides dimidiatus) tending to Ribboned sweetlips (Plectorhinchus polytaenia). Yillet Kecil, Yillet Islands, Misool, Raja Ampat, West Papua, Indonesia
RF - High magnification photo of Secretary blenny (Acanthemblemaria maria) in boulder brain coral (Colpophyllia natans). East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Imitator damselfish (Pomacentrus imitator) male, abstract in in front of a Black coral tree (Antipathes sp.) on a coral reef. Ra Province, Viti Levu, Fiji, Polynesia
Bohar snappers (Lutjanus bohar) breaking up into smaller groups to spawn. The fish swim short arches above the main group as they release clouds of gametes. Shark City, Ulong, Rock Islands, Palau
Portrait of a male River blenny (Salaria fluviatilis) River Flumendosa, Gennargentu National Park, Sardinia, Italy
Rich coral growth, with a pair of Exquiste butterflyfish (Chaetodon austriacus), flourishes beneath the barren desert cliffs in the Red Sea. Taken at sunset. Ras Katy, Sinai, Egypt
California sea lion (Zalophus californianus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath Eureka oil rig, Los Angeles, California, USA, Pacific Ocean
Pattern detail of a Majestic angelfish (Pomacanthus navarchus). Fiabacet Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea
Portrait of a male Stocky anthias (Pseudanthias hypselosoma). Seraya, Tulamben, Bali, Indonesia. Java Sea