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Ruff (Philomachus pugnax) males fighting at lek, Varanger, Norway, May 2016
Common reed bunting (Emberiza schoeniclus) attacking Common cuckoo (Cuculus canorus), female, Braunschweig, Lower Saxony, Germany, May
Whooper swans (Cygnus cygnus) fighting, Finland, April
Cormorant (Phalacrocorax carbo) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, January
San Cristobal giant tortoises (Chelonoidis chatamensis), fighting, Galapaguera, San Cristobal Island, Galapagos
Two young Wild Boar (Sus scrofa) play fighting in snow. The Netherlands, January
Spotted nutcrackers (Nucifraga caryocatactes) fighting in snow, Vitosha Mountain, Sofia, Bulgaria
White-rumped snowfinch (Montifringilla taczanowskii) two fighting, Tibetan Plateau, Qinghai, China
RF - Red deer (Cervus elaphus) stags fighting, Holkham Park, Norfolk, England, UK, September
Raggy scorpionfish (Scorpaenopsis venosa) disputing territory. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Northerns (Accipiter gentilis) fighting over squirrel carcass, Finland, March MarchNortherns (Accipiter gentilis) fighting over squirrel carcass, Finland, March.. March
Black Grouse (Tetrao tetrix) males fighting in winter, Tver, Russia. April
Blackbuck (Antilope cervicapra) two males, stand off at sunrise, Rajasthan, India
Blackbuck (Antilope cervicapra) two males fighting, Rajasthan, India
Two Ibex (Capra ibex) fighting with baby watching, Triglav National Park, Julian Alps, Slovenia, July 2009
Two European brown bears (Ursus arctos) fighting, Kuhmo, Finland, July 2009
Northern gannets (Morus bassanus) fighting, Saltee Islands, Ireland, May 2008
Northern gannets (Morus bassanus) on rock squabbling, Saltee Islands, Ireland, June 2009
Two European bison (Bison bonasus) fighting, Bialowieza NP, Poland, February 2009
Two Reindeer (Rangifer tarandus) fighting, Forollhogna National Park, Norway, September 2008
Two Common starlings (Sturnus vulgaris) fighting, Pusztaszer, Hungary, May 2008. Magic Moments book plate
Two European bison (Bison bonasus) fighting, Bialowieza NP, Poland, February 2009. WWE INDOOR EXHIBITION
Lammergeier / Bearded vulture (Gypaetus barbatos) adult and juvenile squabbling over food in snow, Cebollar, Torla, Aragon, Spain, November 2008. WWE INDOOR EXHIBITION
Two Grey seals (Halichoerus grypus) fighting, Donna Nook, Lincolnshire, UK, November 2008
Reindeer (Rangifer tarandus) fighting, Forollhogna National Park, Norway, September 2008 WWE BOOK
Black tailed godwits (Limosa Limosa) fighting, Lake Belau, Moldova, June
Two Konik horse stallions fighting during breeding season, Oostvaardersplassen, Netherlands, June 2009. WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS
Konik horse, stallions squaring up ready to fight, Oostvaardersplassen, Netherlands, June 2009
Grey seals (Halichoerus grypus) fighting, Donna Nook, Lincolnshire, UK, November 2008
Chaffinch (Fringilla coelebs) two males squabbling at water, Pusztaszer, Hungary, May 2008
Two Eurasian brown bear (Ursus arctos) cubs fighting, Suomussalmi, Finland, July 2008
Alpine ibex (Capra ibex ibex) fighting, Hohe Tauern National Park, Austria, July 2008
Alpine ibex (Capra ibex ibex) fighting in front of a glacier, Hohe Tauern National Park, Austria, July 2008
European Hares (Lepus europaeus) boxing, female on right. Wales, UK, June. Sequence 1 of 2
Two Fallow deer (Dama dama) bucks during fight, rutting season, Richmond Park, London, UK, October
Black grouse (Tetrao tetrix) males fighting at lek site, Cairngorms National Park, Grampian, Scotland, UK, April
Two Grey herons (Ardea cinerea) squabbling over fish fed to them by visitors, River Tame, Reddish Vale Country Park, Stockport, Greater Manchester, UK, December 2010
Three Grey herons (Ardea cinerea) fighting in Regents Park, London, UK, April 2011
Grey seals (Halichoerus grypus) two adults fighting in the surf, Donna Nook, Lincolnshire, England, UK, October
Two common buzzards (Buteo buteo) fighting on ground in snow, Scotland, UK, January
Two Black grouse (tetrao tetrix) males fighting at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Musk ox (Ovibos moschatus) portrait, Wrangel Island, Far Eastern Russia, March
Two male Alpine ibex (Capra ibex ibex) fighting in front of the Mer de Glace glacier, with a female watching, Aiguilles Rouges (Red Peaks) Regional Natural Park, Haute-Savoie, France, June
Male Panther Chameleons (Furcifer pardalis) fighting. Nosy Be, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, January 2010
Bengal Tigers (Panthera tigris) sub-adults, approximately 17-19 months old, playfighting. Endangered. Bandhavgarh National Park, India. Non-ex. Sequence 1 of 2
Red foxes (Vulpes vulpes) fighting in snowy field with woodland in the background, Vosges, France, February
Two Gannets (Morus bassanus) squabbling, watched by another in the background. Hermaness National Nature reserve, Shetland Isles, Scotland
African Buffalo (Syncerus caffer) males fighting, Nakuru National park, Kenya
