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Two Western grey plantain-eaters (Crinifer piscator) looking at each other, Allahein River, The Gambia
Two male Black grouse (Tetrao tetrix) fighting for territory on lek, Cairngorms National Park, Scotland, UK. May
Two male Polar bears (Ursus maritimus) standing on hind legs, sparring, Churchill, Canada. November
Two male Alpine Ibex (Capra ibex) fighting on a mountain slope at dawn, Varaita Valley, Alps, Italy, July
Two Great white egret (Ardea alba) in winter, Kiskunsag National Park, Hungary. December
Grey heron (Ardea cinerea) and Great white egret (Ardea alba) standing facing each other at sunrise. Lake Csaj, Kiskunsagi National Park, Hungary. December
Pair of Great crested grebes (Podiceps cristatus) performing the weed dance during courtship, The Netherlands, Europe. March
Two male Darjeeling bush frogs (Raorchestes Annandalii) fighting over territory, Kalimpong, West Bengal, India
Black stork (Ciconia nigra) and Grey heron (Ardea cinerea) Pusztaszer reserve, Hungary
RF - King penguins (Aptenodytes patagonicus) on the beach at Salisbury Plain. South Georgia, South Atlantic. (digitally stitched image)
RF - Herd of American Bison (Bison bison) around geo-thermal features. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January 2008
Male Anole Lizards (Anolis sp) in territorial display to each other. Rainforest canopy, Rio Claro Reserve, Magdalena Valley, Colombia, South America
Two wild Mustang stallions fighting in White Mountain Herd Area, Wyoming, USA. August
RF- Two Small red damselflies (Ceriagrion tenellum) covered in morning dew. Arne (RSPB) Nature Reserve, Dorset, UK. August
Black necked grebe (Podiceps nigricollis) pair at night, performing their courtship dance during the mating season, with street lights reflected in the water. The Netherlands.April 2014
Arctic foxes (Vulpes lagopus) juveniles playing, in summer coat, Wrangel Island, Far Eastern Russia, August
Musk ox (Ovibos moschatus) portrait, Wrangel Island, Far Eastern Russia, March
Hooded crow (Corvus coronix) and Eurasian Magpie (Pica pica) waiting to scavenging White tailed eagles (Haliaeetus albicilla) food, Pusztazer, Hungary
Great reed warbler (Arcocephalus arundinaceus) chick begging for food, with bokeh affect from water, Pusztaszer, Hungary, June
Three Silver studded blue butterflies (Plebejus argus) covered in dew, Klein Schietveld, Brasschaat, Belgium
Wild Mustang foals among wild flowers, Pryor Mountains, Montana, USA
Silver-studded blue butterfly (Plebejus argus) pair mating, resting on a Pyrimidal orchid (Anacamptis pyramidalis) flower, Gwithian Towans, Cornwall, UK, July
Ringtailed Lemurs playing (Lemur catta) Nahampoana Reserve, South Madagascar, Africa
Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April
Impala (Aepyceros melampus) females, Masai-Mara Game Reserve, Kenya
Chimpanzee (Pan troglodytes) mother and baby age two and a half, tropical forest, Western Uganda
Blue Sheep (Pseudois nayaur) on hillside, Kekexili, Qinghai, China, January
Bengal Tigers (Panthera tigris) sub-adults, approximately 17-19 months old, playfighting. Endangered. Bandhavgarh National Park, India. Non-ex. Sequence 1 of 2
Bengal Tiger (Panthera tigris tigris) mother and cub, Pench National Park, Madhya Pradesh, India, taken on location for Tiger - Spy in the Jungle
Great crested grebe (Podiceps cristatus) pair performing courtship display at dawn, backlit with surrounding by mist, Cheshire, UK, March
RF- Puffins (Fratercula) displaying. Skomer Island, Wales, UK. April
Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June
Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February
Two male Mute swans (Cygnus olor) displaying on water in territorial dispute, UK
Two Denise's Pygmy Seahorses (Hippocampus denise) on a seafan (Muricella sp. ). Digital compositeTwo Denise's Pygmy Seahorses (Hippocampus denise) on a seafan (Muricella sp.). Digital composite. Misool, Raja Ampat, West Papua, Indonesia, November
Western Cattle Egret (Bubulcus ibis) in a field by domestic cattle. River Allier, France, May
Two orphan baby Elephants (Loxodonta africana) being affectionate. David Sheldrick Wildlife Trust Nairobi Elephant Nursery, Kenya, July 2010
Feral domestic horses (Equus caballus) wild herd stallion (grey) fighting a bachelor chestnut stallion, Blue Swallow Reserve, near Kaapsehoop, Mpumalanga province / Eastern Transvaal, South Africa
European hare (Lepus europaeus) mating pair boxing in field, Slovakia
These hatchlings of an Olive ridley sea turtle (Lepidochelys olivacea) should be buried and protected under 40 cm of warm sand, but a domestic dog dug them out at Ostional beach in Costa Rica
Two Mute swan (Cygnus olor) cygnets, Dorset, UK
Two male Thomsons gazelles (Eudorcas thomsonii) fighting, Masai-Mara game reserve, Kenya
Confrontation between a great egret (Ardea alba) and grey heron (Ardea cinerea), Lake Csaj, Kiskunsagi National Park, Hungary
