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Mashpi torrenteer frog (Hyloscirtus mashpi) juvenile. resting on a leaf at night, Mashpi, Pichincha, Ecuador. Endangered
Rufescent tiger heron (Tigrisoma lineatum) portrait, Yasuni National Park, Orellana, Ecuador
Black-headed cat snake (Boiga nigriceps) coiled, portrait, Kubah National Park, Sarawak, Malaysia
Black-mantled tamarin (Leontocebus nigricollis) sitting on branch in rainforest, Cuyabeno Reserve, Sucumbios, Ecuador
Puma (Puma concolor) female with cub, resting, Torres del Paine National Park, Magallanes, Chile
Sri Lankan leopard (Panthera pardus kotiya) resting on tree branch, Yala National Park, Southern and Uva Provinces, Sri Lanka
Sally Lightfoot crab (Grapsus grapsus) on beach, portrait, Santiago Island, Galapagos National Park, Galapagos Islands
Two Puma (Puma concolor) cubs playing, feet pads visible, Torres del Paine National Park, Magallanes, Chile
Ecuadorian squirrel monkey (Saimiri cassiquiarensis macrodon) portrait, Yasuni National Park, Orellana, Ecuador
Red fox (Vulpes vulpes) in snow-covered agricultural field, Bialowieza National Park, Poland. January
Two Pallas's cat (Otocolobus manul) kittens waiting at their den for their mother to return from hunting, Mongolia, June, 2017. Filmed for BBC series Big Cats
A female Pallas's cat (Otocolobus manul) with two young kittens hiding amongst rocks at their den site, Mongolia, June, 2017. Filmed for BBC series Big Cats
Pair of Japanese dwarf flying squirrels (Pteromys volans orii) during the reproductive season, on branch nlooking down, Hokkaido, Japan. The female is on the left, male right
Verreaux's sifaka (Propithecus verreauxi) sitting on log in forest, Berenty forest, southern Madagascar. Critically endangered
Red-fronted brown lemur (Eulemur rufus) female, portrait, Kirindy Forest, Madagascar
White-tailed deer (Odocoileus virginianus) fawn, standing among wildflowers and long grass, Acadia National Park, Maine, USA. June
Raccoon (Procyon lotor) drinking from a beaver pond in Acadia National Park, Maine, USA. June
Atlantic puffin (Fratercula arctica) portrait, Machias Seal Island, Maine, USA. July
Black grouse (Lyrurus tetrix) male, jumping, part of a dominance display, at lek in early morning light, Kuusamo, Finland, April
Longhorn cowfish (Lactoria cornuta) juvenile, swimming over seabed at night, Bitung, North Sulawesi, Indonesia, Molucca Sea
Silvertip shark (Carcharhinus albimarginatus) portrait, Revillagigedo Islands, Mexico, Pacific Ocean
Yellowmargin moray (Gymnothorax flavimarginatus) peering out from Digitate leather coral (Sinularia sp)Yellowmargin moray (Gymnothorax flavimarginatus) peering out from Digitate leather coral (Sinularia sp.) on a reef, Ari Atoll, Maldives, Indian Ocean
Sabre squirrelfish (Sargocentron spiniferum) sheltering beneath a large Table coral (Acropora sp.) on a coral reef, Laamu Atoll, Maldives, Indian Ocean
Leopard (Panthera pardus) hiding in dense foliage, Bardia National Park, Terai, Nepal
Bengal tiger (Panthera tigris tigris) walking through river, snarling, Bardia National Park, Terai, Nepal. Endangered
Changeable hawk-eagle / Crested hawk-eagle (Nisaetus cirrhatus) perched on branch, Bardia National Park, Terai, Nepal
Grey langurs (Semnopithecus entellus) family group, sitting and climbing in tree, Bardia National Park, Terai, Nepal
Koala (Phascolarctos cinereus) resting during the day in large Eucalypt (Eucalyptus sp)Koala (Phascolarctos cinereus) resting during the day in large Eucalypt (Eucalyptus sp.) tree, Goonderoo Nature Reserve, Queensland, Australia
Frog (Anura) in waterhole at night with two Cape buffalo (Syncerus caffer) in background, Zimanga Game Reserve, KwaZulu-Natal, South Africa
Snowy owl (Bubo scandiaca) perched on wodden post at dusk, Canada. February
Snowy owl (Bubo scandiaca) standing behind dried plant in snow, Canada. January
Female Arctic fox (Vulpes lagopus) in snow, Hornstrandir, Iceland. Blue colour morph in winter coat. February
Grey-headed flying-fox bat (Pteropus poliocephalus) hanging from branch looking down with wing extended, Yarra Bend Park, Victoria, Australia. Cropped
Great horned owl (Bubo virginianus) juvenile, perched on branch, Texas, USA. May
American bobcat (Lynx rufus) male, drinking at water's edge, Texas, USA. April
Close up of Galapagos hawk (Buteo galapagoensis) in immature plumage, flying, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
Snowy owl (Bubo scandiaca) female, head portrait, Canada. January
Arctic fox (Vulpes lagopus), female blue colour morph in winter coat, during snow blizzard. Baited with small amount of natural food (dried fish). Hornstrandir, Iceland. February
Mountain hare (Lepus timidus) with fur encrusted in snow, Cairngorms National Park, Scotland, UK. January
Adelie penguin (Pygoscelis adeliae) leaping out of ocean as it approaches land, Devil Island, Antarctica, Southern Ocean
Lioness (Panthera leo) lying in long grass on the savannah, portrait, Okavango Delta, Botswana, Africa
Lion (Panthera leo) cub, aged 3 months, climbing on branch and chewing on a stick, Okavango Delta, Botswana, Africa
Two Dwarf mongoose (Helogale parvula) peering out of their burrow in termite mound, Okavango Delta, Botswana, Africa
Four Common / Plains Zebras (Equus quagga) standing side by side, portrait, Okavango Delta, Botswana, Africa
Painted frogfish (Antennarius pictus) portrait, Sipidan, Malaysia, Celebes Sea
Asian small-clawed otter (Aonyx cinerea) pups looking at camera. Captive, occurs in Asia. Zooparc Overloon, the Netherlands
Male Highland Great diving beetle (Dytiscus lapponicus) on surface of water, Darnaway Estate, Highlands, Scotland, UK. May. Focus stacked. Captive
Kerry bog pony, gelding, a rare breed, head portrait, County Kerry, Republic of Ireland. April
