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Alpine ibex (Capra ibex) female, Jura, Switzerland, September
Great egret (Ardea alba) two fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Alpine ibex (Capra ibex), male in habitat, Bernese Alps, Switzerland. July
Bearded tit (Panurus biarmicus) male portrait feeding, Helsinki, Finland, January
Sgwd Ddwli Uchaf, Brecon Beacons National Park, Wales. June 2013
Pebbles on Nash Point, Monknash Coast, Vale of Glamorgan, Wales, UK, June 2006
Aerial view of the East Lighthouse, Rathlin Island, County Antrim, Northern Ireland, UK, September 2009
RF- White horse of the Camargue, running from sea. Camargue, Southern France
Candover Brook, tributary of the River Itchen, chalk stream Hampshire, England, UK, June. Photographed for The Freshwater Project
Male Greenfinch (Carduelis chloris) amongst Pussy willow catkins, Hertfordshire, England, UK
Male Bullfinch (Pyrrhula pyrrhula) perched amongst Cherry blossom, Buckinghamshire, England
Fieldfare (Turdus pilaris) in Rowan tree with berries in winter, Jyvaskyla, Central Finland, January
Oystercatcher (Haematopus ostralegus) by the coast, Island of Runde, Norway July
Mosquito (Aedes punctor) female resting after sucking blood from human arm. Sequence 4 / 4Mosquito (Aedes punctor) female resting after sucking blood from human arm. Sequence 4/4
Two Arctic hares (Lepus arcticus) fighting, Northeast Greenland National Park, Greenland. Veolia Environnement Wildlife Photographer of the Year 2009: Mammal Behaviour category: Runner-up
Two Edelweiss (Leontopodium alpinum) flowers, Triglav National Park, Slovenia, July 2009
Mount Prisojnik (2, 547m) and Mount Razor (2, 601m), viewed from Kranjska Gora. Triglav National Park, Julian Alps, Slovenia, July 2009
European Hare (Lepus europaeus) leveret feeding on grass. Wales, UK, May
Little Owl (Athene noctua) taking flight from a post. Wales, UK, June
Oxeye Daisy (Leucanthemum vulgare) flower head at dusk. Scotland, UK, June
Oystercatchers (Haematopus ostralegus) arriving to roost at Snettisham on the Wash. Norfolk, January
Pink-footed Geese (Anser brachyrhynchus) in flight at dusk. The Wash. Norfolk, January
Bearded reedling / tit (Panurus biarmicus) male perched on twig, Titchwell, Norfolk, UK, June
Aerial view over Glenfeshie in winter, Cairngorms National Park, Scotland, UK, January 2012
Coal tit (Periparus ater) adult perched in winter, Scotland, UK, December. Did you know? Coal tits keep in contact with their flock when searching for food, with constant dee or seesee calls
Assynt mountains, Highland, Scotland, UK, June 2011
Star trails above Scots pine (Pinus sylvestris) woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, March 2017
Scots pine (Pinus sylvestris) trees reflected in Loch Maree at dawn with Slioch in background, Wester Ross, Scotland, UK, November 2014
Sunset over Loch Coire Mhic Fhearchair from Beinn Eighe with the sky reflected in the water. Torridon, Highlands, Scotland, UK, June 2016
Pine marten (Martes martes) climbing Scots pine tree, Scotland, UK. May
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Black-headed gull (Chroicocephalus ridibundus) in flight, artistically blurred photograph, Cheshire, UK, February. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
The River Thames and Houses of Parliament at Westminster at sunset with gull, UK, November 2005
Bamburgh Castle and sand dunes in warm, late evening light with stormy evening sky, Bamburgh, Northumberland, UK. March 2014
River Plym flowing fast through Dewerstone Wood, Shaugh Prior, Dartmoor National Park, Devon, England, UK, October 2011. Did you know? The name Plym comes from the Old English term for plum tree
Pine marten (Martes martes) portrait, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Portrait category of the BWPA (British Wildlife Photography Awards) Competition 2016
Hazel Dormouse (Muscardinus avellanarius) hibernating amongst leaves. Captive. Leicestershire, UK, October
Two Rooks (Corvus frugilegus) silhouetted as they perch on a tree branch at their roost site at sunset, Gloucestershire, UK, February
Badger (Meles meles) adult, backlit at sunset, scratching its ears, Derbyshire, UK, May. Winner of the Animal Behaviour category of the British Wildlife Photography Awards (BWPA) Competition 2017
Red deer stag (Cervus elaphus) in heavy snow, Cheshire, UK, March
Red Fox (Vulpes vulpes) cub in late evening light, Leicestershire, England, UK, July
Atlantic Puffin (Fratercula arctica) one landing among resting group, Sule Skerry, Scotland, UK, July
Sand lizard male {Lacerta agilis} Ainsdale NNR, Lancashire, UK
Gannet (Morus bassanus) portrait, Bass Rock, Firth of Forth, Scotland, UK, June
Chlorophyte or green alga (Volvox aureus) in pond water. UK. The larger spheres are approximately 0.5mm in diameter. Digital composite image
Gnarled trunk of an Olive tree (Olea europea) Kolimvaro, Crete, Greece, April 2009
Wolf (Canis lupus) pack running in deep snow, captive in enclosure of the Bavarian Forest National Park, Germany, February