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Wolf (Canis lupus) pack running in deep snow, captive in enclosure of the Bavarian Forest National Park, Germany, February
Twin Birch (Betula) trees highlighted snow. Senja, Norway, February
Portrait of a grey Andalusian stallion, in Ciutadella, Menorca, Spain
Female Scottish wildcat (Felis silvestris grampia) portrait, in winter coat, semi habituated living wild in native oakwood, Lochaber, Scotland, April
Atlantic puffin (Fratercula arctica) gathering around burrows on cliff top, Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland, UK, June
Kilchurn Castle on Loch Awe, Argyll and Bute, Strathclyde, Scotland, UK, April 2011
Adult Adder (Vipera berus) basking on pebble beach. Abbotsbury, Dorset, UK, March
Storm clouds building up off coasts of Norfolk, Weybourne Cliffs, Norfolk, UK, November
Colourful water-smoothed pebbles on shingle beach. Normandy, France, October 2010
Suffolk Punch heavy horse in field resting head anothers back, UK, September
Silhouette of Barn Swallow (Hirundo rustica) flying over water, hawking for insects, Berwickshire, Scotland, UK, July
Split level landscape in oxbow of the Aare river, Switzerland, September
European Lynx (Lynx lynx) adult male stalking through winter birch forest, Bardu, Norway, captive
Herb robert (Geranium robertianum) flower, Coombe Valley, Cornwall, UK, May
Beech (Fagus sylvatica) woodland in spring, at dawn, Peak District National Park, Cheshire, UK, May
Track through fir forest in autumn, Vosges mountain, France, October
View looking west from the summit of Conwy Mountain, with Alltwen Mountain above the farm and Penmaenmawr Mountain in the distance, North Wales, UK, August 2017
Atlantic Puffins (Fratercula arctica) pair bill rubbing, part of ritual courtship, Skomer Island, Wales, UK, May
Mute swan (Cygnus olor) in early morning light removing its own feathers, during moulting, Valkenhorst Nature Reserve, The Netherlands, Europe. August
Red Fox (Vulpes vulpes) walking along a fallen trunk in misty forest. Black Forest, Germany, November
Waterfall on Annalong River, flowing through Mourne Mountains. County Down, Northern Ireland, UK. June 2020
Early purple orchids {Orchis mascula} in flower, Cressbrook Dale, Peak District National Park, Derbyshire, UK, May 2009
Bearded reedling (Panurus biarmicus) female feeding on reed seeds covered in frost, Espoo, Finland, January
White-tailed eagle (Haliaeetus albicilla) taking off with fish prey, Norway, August
Species rich alpine meadow with Orange lily (Lilium bulbiferum), Meadow clary (Salvia pratensis) and Yellow rattle (Rhinathus sp)
Grey heron (Ardea cinerea) wading at night, reflected in water. Lincolnshire, UK, September
RF - White-legged damselfly (Platycnemis pennipes) peering over Oxeye daisy (Leucanthemum vulgare) flower. The Netherlands. August
Fallow deer (Cervus dama) stags at sunset, Holkham Park, North Norfolk, UK, January
Red fox (Vulpes vulpes) in snowy landscape with trees and two Crows (Corfus corone) winter, Jura, Switzerland
Wren (Troglodytes troglodytes) singing, early morning in spring, Broxwater, Cornwall, UK. April
Common oak (Quercus robur) fowers shedding pollen. Surrey, UK. April
Small Tortoiseshell Butterfly (Aglais urticae) feeding on scabious flowers in hay meadow, Wensum Valley, Norfolk, England, UK, July
Buckholt Wood is a beech (Fagus sylvatica) wood and part of the Cotwolds Commons and Beechwoods National Nature Reserve, on the scarp slope of the Cotswolds near Birdlip, Gloucestershire
Red Deer stag and hind (Cervus elaphus) silhouetted at sunset, Holkham Park, Norfolk, UK, December
Goshawk (Accipiter gentilis) plucking a Willow Grouse (Lagopus lagopus) Liminka, Finland, March
Scottish primrose (Primula scotica) native only on the northern coast of Scotland and Orkney, here growing on grasslands at Dunnet Head, Caithness, England, UK, July
Early or English gentian (Gentianella anglica) a scarce plant, endemic to England, growing here at Durlston Head in Dorset, England, UK, May
Mother of pearl (Pleuroptya ruralis) moth nectaring on Ragwort (Jacobaea vulgaris) at night. Surrey, England, UK. July
Garden bumblebee (Bombus hortorum) queen nectaring on Comfrey (Symphytum Hidcote Pink ) flowers. In garden, Surrey, England, UK. March
Sanderling (Calidris alba) flock foraging at sunset, Britanny, France, October
Peregrine Falcon (Falco peregrinus) perched on a cliff above a woodland in autumn. Kinnoull Hill Woodland Park, Perthshire, Scotland, November
Scottish wildcat (Felis silvestris grampia) captive, endemic to Scotland. September
Shafts of light at dusk over Wrynose pass and the Coniston hills from Ambleside, Lake District, England, UK. September 2015
Rocky shoreline, Loch Scavaig and view to Cuillin mountians, Isle of Skye, Scotland, UK, April
Village Bay on St Kilda and the old house that formed the village, Scotland, UK, May
Red squirrel (Sciurus vulgaris) sitting on fallen tree in pine woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, August
Gannet (Morus bassanus) fluffy chick with its parent. Shetland Islands, Scotland, UK, August
Small tortoiseshell butterfly (Aglias urticae) on thistle. Dorset, UK, August