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Young Polar bear (Ursus maritimus) walking across melting sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Blacktip reef sharks (Carcharhinus melanopterus) swimming in shallow crystal clear water, Aldabra Atoll, Seychelles, Indian Ocean
Massing herds of White bearded wildebeest (Connochaetes taurinus albojubatus) on migration, Serengeti National Park, Tanzania
Aerial view of two Gemsbok (Oryx gazella) in sand dunes, Namib Desert, Namibia
Nile crocodile (Crocodylus niloticus) making its way through sand to the Rufiji River, Selous Game Reserve, Tanzania
Hraunfossar waterfall in autumn, Iceland, September 2013
Wildebeest (Connochaetes taurinus) crossing the Mara river, Masai-Mara Game Reserve, Kenya
Great Blue shark (Prionace glauca) viewed from above, Pico Island, Azores, Portugal, Atlantic Ocean
Mineral rich landscape and copper lake from the Bogo La pass at17000 ft, Tibet, June 2010
Rio Jatuntianhua Robber Frog (Pristimantis eriphus) male on a leaf. Ecuador, Vulnerable species
Newlands Valley and Causey Pike, Lake District National Park, Cumbria, England, UK. October 2012
Lammergeier / Bearded Vulture (Gypaetus barbatus) in flight. Ordesa National Park, Spanish Pyrenees, April
Tibetan antelope (Pantholops hodgsonii) herd in the early morning with a low moon, Kekexili, Qinghai, Tibetan plateau, China, January
Nigel Creek with conifer forest at dusk, Banff National Park, Rocky Mountains, Alberta, Canada
Domestic cows (Bos taurus) and free roaming horses grazing in the Pyrenees-Atlantiques, Pyrenees, France, June
Aerial view of the Long Wall, sand dunes along the Atlantic coast of the Namib desert, Swakopmund, Namibia, August 2008
Aerial view saltmarshes of Bahia de Cadiz Natural Park, Huelva, Andalucia, Spain, March 2008
Aerial image showing sandbanks in the Bahamas archipelago, Caribbean, February 2012
Aerial view of Everglades National Park, Florida, USA, February 2012
Northern lights / Aurora borealis over snowy landscape, Iceland, Europe, March 2012
Sand dunes in the Libyan desert, Sahara, Libya, North Africa, December 2007
Lake Ann surrounded by Douglas firs (Pseudotsuga menziesii) Silver firs (Abies alba) and other conifers in the area of Rainy Pass of North Cascades National Park, North Cascades National Park
Aerial view of Grand Prismatic thermal spring in Yellowstone National Park, USA. The bright colours are natural, coming from thermophile bacteria in the water
Tench (Tinca tinca) on the surface of a turbid marsh pond, with Rudd (Scardinius erythrophthalmus) above, Fribourg, Switzerland, 2004
A small valley in Alsace seen from a high angle. France
A thunderstorm drops heavy rain over the hoodoo sandstone formations. Bryce Canyon National Park, Utah, USA, August
Aerial view of the rivers and saltmarshes of the Bahia / Bay de Cadiz Natural Park, Andalucia, Spain, March 2008
Fogbow over pack ice, Svalbard, Arctic Norway 2010
A lone Caribbean reef shark (Carcharhinus perezi) cruises over sand ripples. Walkers Cay, Northern Bahama Islands, Republic of Bahamas
Rain shower approaching La Vieille lighthouse, Pointe du Raz, Brittany, France. November 2006
Wild flower meadow in city centre, Sheffield, Yorkshire, UK, August 2009
Aerial view of Hidden Bay in North West peninsula of Gam Island. Raja Ampat Islands, West Papua, Indonesia
RF - Lowland rainforest, Osa Peninsula, Costa Rica
Aerial view of Delta Sierpe River Terraba, Corcovado National Park, Osa Peninsula, Puntarenas Province, Costa Rica. December 2014
Dawn over the Segama River, with mist hanging over lowland rainforest. Heart of Danum Valley, Sabah, Borneo
Tail fluke of a diving Humpback whale (Megaptera novaeangliae) misty coast in background, Alaska, USA. Gulf of Alaska, Pacific ocean
Fulmar (Fulmarus glacialis) rear view of bird hanging in air over steep cliffs, Shetland Islands, Scotland, UK. HIGHLY COMMENDED - Animals in their environment - Wildlife Photographer of the Year
Common dolphin (Delphinus delphis) surfacing, Atlantic ocean, Portugal, September
Aerial view of Humpback whales (Megaptera novaeangliae) Bay of Fundy, Canada. November
Lomnicks Peak, 26345m, one of the highest mountain peaks in the High Tatras mountains of Slovakia June 2012
Andean Flamingoes (Phoenicoparrus andinus) on the Altiplano, Sur Lipez Province, Potosi Department, Bolivia. December 2013
Succulent (Aeonium pseudourbicum) close up covered in dew, Tenerife, Canary Islands, Europe
Mist hanging over Lowland Dipterocarp Rainforest just after sunrise. Danum Valley Conservation Area, Sabah, Borneo
Aerial panorama of Mounu Island Resort in Vava'u, Tonga. September 2017
Kittiwakes (Rissa tridactyla) flying over surf, Vardo, Norway March
Green sea turtle (Chelonia mydas) swimming over sand ripples in shallow water, Eleuthera, Bahamas. Endangered
Iberian ibex (Capra pyrenaica), pair with male top right, during rutting season. Sierra Nevada National Park, Andalusia, Spain. October