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View of Yamdrok-Tso Lake (at 4, 441m), sacred for Tibetans. Tibet, China. August, 2015
Prayer flags on shores of Nam-Tso Lake. Tibet, China
Domestic yak (Bos grunniens) lying under prayer flags near shores of Nam-Tso Lake (at 4, 730m), second largest salt lake in China. Tibet, China
Hundreds of prayer flags at Dolma La Pass, highest point on pilgrimage around sacred Mount Kailash. Tibet, China
Two Pallas's cat (Otocolobus manul) kittens waiting at their den for their mother to return from hunting, Mongolia, June, 2017. Filmed for BBC series Big Cats
A female Pallas's cat (Otocolobus manul) with two young kittens hiding amongst rocks at their den site, Mongolia, June, 2017. Filmed for BBC series Big Cats
Pair of Broadclub cuttlefish (Sepia latimanus) mating in a coral reef, Kagoshima Prefecture, Japan. Having succeeded in gaining the female's acceptance
Female Siberian flying squirrel (Pteromys volans orii) sitting at entrance to nest. Hokkaido, Japan. February
Male Siberian flying squirrel (Pteromys volans orii) attempting to enter nest with female inside during reproductive season but stopped mid-way then rejected. Hokkaido, Japan. March
Composite image showing Siberian flying squirrel (Pteromys volans orii) defecating for duration of about twenty seconds after emerging from nest to forage. Hokkaido, Japan. March
Split-second prior to Siberian flying squirrel (Pteromys volans orii) landing on tree, adjusting its attitude in mid-flight toward relatively vertical position to reduce speed
Leaves of Momiji Japanese maple trees (Acer) turning red in autumn. Kyoto, Japan
Autumn foliage of Momiji Japanese maple trees (Acer). Kyoto, Japan
Pair of Siberian flying squirrel (Pteromys volans orii) sharing nest in tree hole in preparation for reproductive season. Hokkaido, Japan. March
Spawning Leaf corals (Pavona decussata). Spawning is synchronized over a large area with many coral colonies, Kyushu, Japan, Pacific Ocean. July
Reef manta ray (Mobula alfredi) swimming with a Twinspot snapper (Lutjanus bohar), Okinawa, Japan, Pacific Ocean
Female Northern pygmy squid (Idiosepius paradoxus) preparing to lay another egg on a blade of eelgrass (Zostera marina), Yamaguchi Prefecture, Japan
Pair of Japanese dwarf flying squirrels (Pteromys volans orii) during the reproductive season, on branch nlooking down, Hokkaido, Japan. The female is on the left, male right
Long-tailed tit (Aegithalos caudatus) feeding on icicle formed from sap of Painted maple tree (Acer pictum). Hokkaido, Japan. February
Male Korean seahorse (Hippocampus haema) giving birth. Kumamoto Prefecture, Kyushu, Japan
White-tailed eagle (Haliaeetus albicilla) taking off from iceberg at sunrise, Shiretoko National Park, Hokkaido, Japan
Whooper swan (Cygnus cygnus) in flight with buildings behind, Sanmenxia, Henan province, China
Whooper swans (Cygnus cygnus) in flight with cityscape behind, Sanmenxia, Henan province, China
Whooper swans (Cygnus cygnus) at sunset, Sanmenxia, Henan province, China
Whooper swan (Cygnus cygnus) outstretching its wings, Sanmenxia, Henan province, China
Whooper swans (Cygnus cygnus) in flight, Sanmenxia, Henan province, China
Kweichow crocodile newt (Tylototriton kweichowensis) feeding on a worm, China
Autumn leaves, Japanese maple (Acer palmatum) Lu Shan, Taiwan
Sumatran tiger (Panthera tigris sondaica) portrait, captive
Banded houndshark (Triakis scyllium) group above sea floor. Chiba Prefecture, Honshu, Japan. October
White-lipped deer (Cervus albirostris) walking on the Tibetan plateau, Serxu, Garze Prefecture, Sichuan Province, China
Golden Birdwing butterfly (Troides aeacus) Sheding Forest park, Kenting National Park, Taiwan
Cloud forest in the Yushan National Park, Taiwan
Fern forest in the Yushan National Park, Taiwan
Taiwan Bar-wing (Actinodura morrisoniana) an endemic species, feeding on red berry in tree, Lu Shan, Taiwan
Taiwan Yellow tit, (Machlolophus holsti) Lu Shan, Taiwan endemic species
Bamboo, Daxueshan National Recreational Forest, Taiwan
Formosan rock macaque (Macaca cyclopis) social grooming behaviour. Alishan National Recreational Forest, Taiwan
Misty mountains, Alishan National Recreational Forest, Taiwan
Swinhoes pheasant (Lophura swinhoii) male, endemic species, Yunlinshan, Taiwan
Black-eared kite, (Milvus migrans lineatus), Bi Tan Bay, Taiwan
Snowflake moray (Echidna nebulosa) Green Island, Taiwan
Semicircle angelfish (Pomacanthus semicirculatus) Kenting National Park, Kenting, Taiwan
Bargibants seahorse (Hippocampus bargibanti) Kenting Nationalpark, Kenting, Taiwan
Tiny hydrozoa, which is a predatory jellyfish-like animal. The hydrozoans are related to jellyfish and corals and belong to the phylum Cnidaria, Green Island, Taiwan
Juvenile Flying fish (Cypselurus sp), Longdong, Northeast, Taiwan
Red stingray (Hemitrygon akajei) with Banded houndshark (Triakis scyllium) group and smaller fish in background. Chiba Prefecture, Honshu, Japan. September
Red stingray (Hemitrygon akajei), four in a row, viewed from below. Chiba Prefecture, Honshu, Japan. October