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Male Leach's sea star (Leiaster leachi) broadcast spawning, releasing streams of sperm from its arms. Kagoshima Prefecture, Kyushu, Japan
Siberian flying squirrel (Pteromys volans orii) flying directly over photographer. Hokkaido, Japan. February
Indo-Pacific bottlenose dolphin (Tursiops aduncus) with penis extended. Ogasawara / Bonin Islands, Japan
Portrait of a Golden snub-nosed monkey (Rhinopithecus roxellana) screaming and showing its teeth, Foping Nature Reserve, Shaanxi, China. Endangered species
Hong Kong orchid tree (Bauhinia blakeana) flowers. Hong Kong, December
Stellers sea-eagle (Haliaeetus pelagicus) in flight, Hokkaido, Japan, February
Japanese macaque (Macaca fuscata) sleeping at hot spring in Jigokudani, Yaenkoen, Nagano, Japan, February
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
Blue poppy (Meconopsis zangnanensis) Mt Qomolangma National Park, Qinghai Tibet Plateau, China
Whooper swans (Cygnus cygnus) three coming into land, Hokkaido, Japan, February
Snow leopard (Uncia uncia) Altai Mountains, Mongolia. March
Japanese cranes (Grus japonensis) displaying in snow, Hokkiado, Japan, February
Stellers sea-eagle (Haliaeetus pelagicus) landing on pack ice, Hokkaido, Japan, February
Red panda (Ailurus fulgens) captive, occurs in China
Japanese Macaques (Macaca fuscata) in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Eurasian crag martin (Ptyonoprogne rupestris) Kawakarpo Mountain, Meri Snow Mountain National Park, Yunnan Province, China. May
Kiang (Equus kiang) herd grazing on steppe in evening light, mountains in background. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October 2019
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Backlit photo of a Robust ghost pipefish (Solenostomus cyanopterus) reveals a small fish that it has just eaten in its stomach. Lembeh Strait, North Sulawesi, Indonesia. Molluca Sea
Three Red-crowned cranes (Grus japonensis) in flight. Hokkaido, Japan. March
Mimika bobtail squid (Euprymna morsei), Hokkaido, Japan, Pacific Ocean
Lady Amhersts pheasant (Chrysolophus amherstiae) Kanding, Sichuan, China
Proboscis monkey (Nasalis larvatus) male, Sabah, Borneo
Leaves of a Ginkgo tree or Maidenhair tree (Ginkgo biloba) Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Japanese Rhinoceros beetle (Allomyrina dichotoma dichotoma) male, Guangshui, Hubei province, China. July
Firefly squid (Watasenia scintillans) emitting light from photophores, Toyama Bay, Japan. This form of bioluminescence is used as counter illumination camouflage
Female Pallass cat (Otocolobus manul) suckling four young kittens at their den, Mongolia, June, 2017. Filmed for BBC series Big Cats
Large leaf of Giant Elephants Ear (Alocasia odora), in rainforest, Yangminshan, Taiwan
Osprey (Pandion haliaetus) carrying fish above river in Taipei, Taiwan
Tibetan gazelle / Goa (Procapra picticaudata) Keke Xili / Hoh Xil Nature Reserve, Tibetan High plateau, Qinghai, China
Mandarin duck (Aix galericulata) male female pair, Yuyuantan Park, Beijing, China
Golden snub-nosed monkey (Rhinopithecus roxellana), mother and baby, Qinling Mountains, Shaanxi province, China
Altai argali sheep (Ovis ammon), three grazing on slope. Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China. October
Tibetan gazelle (Procapra picticaudata) in snow. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October
Pallass cat (Otocolobus manul) licking lips whilst running over ice. East Mongolia. February
Lady Amhersts pheasant (Chrysolophus amherstiae) male displaying, Kanding, Sichuan, China. April
Siberian flying squirrel (Pteromys volans orii) feeding on Sakhalin fir tree. Hokkaido, Japan. March
Asian sheepshead wrasse (Semicossyphus reticulatus) mature male in breeding colouration. Niigata, Japan
Sperm whale (Physeter macrocephalus) female at the ocean surface with her mouth open. This whale has just surfaced from foraging in deep water. Ogasawara or Bonin Islands, Japan
Giant panda (Ailuropoda melanoleuca) cub in tree, Chengdu Panda Breeding Centre, Sichuan, China. Captive
Siberian weasel (Mustela sibirica) standing in snow. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Sesame (Sesamum indicum), green unripe pods. Sichuan Province, China
Hummingbird hawk-moth (Macroglossum stellatarum) nectaring on Loofah (Luffa sp) flower. Xinjiang, China
Mandarin duck (Aix galericulata) pair mating on lake. Shanghai Zoo, China
Shrubs, trees and lianas in tropical rainforest. Xishuangbanna, Yunnan Province, China
Euphorbia (Euphorbia jolkinii) leaves turning red in autumn. Napa Hai, Shangri-la, Yunnan Province, China. September
Cut bamboo stems stacked in a pile, end on. Anhui Province, China
Chinese grass carp / White amur (Ctenopharyngodon idella), farmed for food in China. Captive specimen, China