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White rhinoceros (Ceratotherium simum) with unusually shaped horn, Solio Game Reserve, Laikipia, Kenya. September
Dwarf forest chameleon (Rhampholeon boulengeri), Nyungwe Forest National Park, Rwanda
Portrait of male silverback Mountain gorilla (Gorilla beringei beringei) feeding. Bwindi Impenetrable Forest National Park, Uganda, Africa
Great blue turaco (Corythaeola cristata) in Bigodi wetland santuary, Uganda
Olive baboon (Papio hamadryas anubis) mother with babies in a tree. Kibale National Park, Uganda, Africa
Chimpanzee (Pan troglodytes schweinfurthii) male vocalising in tree, Kibale National Park, Uganda, Africa
Mountain gorilla (Gorilla beringei) mother with one month baby feeding in tree. Member of Katwe group. Bwindi Impenetrable Forest National Park, Uganda, Africa
Mountain gorilla (Gorilla berengei), aggressive young male from Titus Group, Visoke, Volcanoes National Park, Rwanda, August
Ruppells griffon vulture (Gyps rueppellii) flying over Jinbar waterfall, Simien Mountains National Park, Amhara, Ethiopia, September. Critically endangered species
Gelada Baboon (Theropithecus gelada) male lying in a forest glade at around 3, 000 metres of altitude, Simien Mountains National Park, Amhara, Ethiopia
Cheetah (Acinonyx jubatus) female and cubs. Brood of seven cubs, a record for the area. Masai Mara National Reserve, Kenya
Cheetah (Acinonyx jubatus) female resting with cubs climbing on back. Masai Mara National Reserve, Kenya
RF - Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) waiting to cross the Mara River. Serengeti National Park, Tanzania. September
Nile crocodile (Crocodylus niloticus) waiting to attack White-bearded wildebeest (Connochaetes taurinus albojubatus) as they cross the Mara River
African elephant (Loxodonta africa) male in threat display to rival. Floor of the Ngorongoro Crater, Ngorongoro Conservation Area, Tanzania
RF - Male African lion (Panthera leo) scratching tree at Big Marsh, Nogorongoro Conservation Area / Serengeti National Park, Tanzania. March
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at dawn. Kasanka National Park, Zambia
Female cheetah (Acinonyx jubatus) named Silgi (means bright future in Swahili) walking with 7 cubs. Masai Mara National Reserve, Kenya
Ruwenzori side-striped chameleon, (Trioceros rudis), hanging from fern, Nyungwe Forest NP, Rwanda, December. Non-ex
Balsam (Impatiens rosulata) with yellow nectar guides. Endemic to Kitulo Plateau, Tanzania
Spiderplant (Cleome hirta) flowerhead. Patterns on two banner petals serve as nectar guidesSpiderplant (Cleome hirta) flowerhead.Patterns on two banner petals serve as nectar guides. Tanzania
Reticulated Giraffe (Giraffa camelopardalis reticulata) standing in grassland. Laikipia, Kenya. October
Chimpanzee juvenile (Pan troglodytes schweinfurthii) in a tree. Kibale National Park, Uganda. January
Chimpanzee male (Pan troglodytes schweinfurthii) sleeping on the forest floor with feet up, Kibale National Park, Uganda. January
Mountain Gorilla (Goriila beringei) portrait, Volcanoes National Park, Virunga Mountains, Rwanda
African hunting dogs (Lycaon pictus) pack mobbing a Spotted hyena (Crocuta crocuta). Hyenas follow the dogs and attempt to steal their kills, so the dogs harass them whenever they get a chance
Swaynes hartebeest (Alcephalus busephalus swaynei), Senkelle Swaynes Hartebeest Sanctuary, Ethiopia
Oribi (Ourebia ourebi montana), Senkelle Swaynes Hartebeest Sanctuary, Ethiopia
Cheetah (Acinonyx jubatus), cubs age 8 weeks playing, Masai Mara Game Reserve, Kenya
Blue wildebeest (Connochaetes taurinus), migrating herd silhouetted on the horizon at sunset, Masai Mara Game Reserve, Kenya
African elephants (Loxodonta africana) grazing on shore of Lake Kariba, Matusadona National Park, Zimbabwe
Katydid (Tettigoniidae) with Giraffe (Giraffa camelopardalis) in the background, Masai Mara National Park, Kenya
Lesser flamingos (Phoeniconaias minor) flock on lake at dawn, Lake Ndutu Tanzania
Reticulated giraffes (Giraffa camelopardalis reticulata) black and white picture of group of three, Kenya
African lion (Panthera leo) male lying in the middle of the Busanga Plains, Kafue National Park, Zambia. Vulnerable
Lionesses (Panthera leo), Masai Mara Game Reserve, Kenya, November
African lioness (Panthera leo) sitting patiently in the long grass, watching herd of Common eland (Tragelaphus oryx). Focus on lion. Grumeti Reserve, Northern Tanzania
Lion (Panthera leo) male resting, Masai Mara Game Reserve, Kenya, November
Lion (Panthera leo) female standing alert, Masai Mara Game Reserve, Kenya
Lioness (Panthera leo) Masai-Mara Game Reserve, Kenya
Black rhino (Diceros bicornis), male on the plains, Masai-Mara Game Reserve, Kenya
Cheetah (Acinonyx jubatus) walking near wildebeest, Masai Mara Nature Reserve, Kenya
Cheetah (Acinonyx jubatus), mother and juvenile cubs, Masai Mara Game Reserve, Kenya
Cheetah (Acinonyx jubatus) male on the hunt with Crater rim in the background, Ngorongoro Crater, Tanzania
Mountain gorilla (Gorilla beringei beringe) female with baby age, 4 months, Volcanoes National Park, Rwanda
Masai giraffe (Giraffa camelopardalis tippelskirchi), female and calf at sunset, with Abdims storks in flight, Masai-Mara Game Reserve, Kenya
Two Lionesses (Panthera leo) trying to defend kill from large pack of Spotted hyaenas (Crocuta crocuta) Masai-Mara game reserve, Kenya
Hippopotamus (Hippopotamus amphibius) male marking territory with faeces, Masai Mara Game Reserve, Kenya