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Laysan albatross (Phoebastria immutabilis), courtship ritual, Eastern island, Midway Atoll National Wildlife Refuge, Hawaii
Arctic grayling (Thymallus arcticus) displaying his vibrant dorsal fin. North Park, Colorado, USA
Ruff (Philomachus pugnax) males fighting at lek, Varanger, Norway, May 2016
Ruff (Philomachus pugnax) male displaying, Varanger, Norway, May 2016
Ruffs displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Ruff displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Capercaillie (Tetrao urogallus) male displaying and calling at lek, Kamchatka, Far east Russia, May
Black Grouse (Tetrao tetrix) lek with male displaying and females around in winter, Tver, Russia. April
Western capercaillie (Tetrao urogallus) Tver, Russia. May
Common Peafowl (Peacock) male displaying, India
Eurasian Spoonbill / White Spoonbill (Platalea leucorodia) in courtship display. Balz, India
Victorias riflebird (Ptiloris victoriae) male displaying in rainforest, Atherton Tablelands, Queensland, Australia, October
Northern gannets (Morus bassanus) displaying, Saltee Islands, Ireland, May 2008
Red deer (Cervus elaphus) stag calling at sunset during rut, Klampenborg Dyrehaven, Denmark, September 2008
White stork (Ciconia ciconia) pair at nest site on old chimney, Rusne, Nemunas Regional Park, Lithuania, June 2009
White stork (Ciconia ciconia) pair silhouetted at nest on old chimney, Rusne, Nemunas Regional Park, Lithuania, June 2009
White stork (Ciconia ciconia) pair displaying, silhouetted at nest on old chimney, Rusne, Nemunas Regional Park, Lithuania, June 2009
White stork (Ciconia ciconia) pair, courtship, at nest on old chimney, Rusne, Nemunas Regional Park, Lithuania, June 2009
Tree sparrow (Passer montanus) displaying in rose bush, Slovakia, Europe, May 2009
RF- Capercaillie (Tetrao urogallus) male displaying in heavy snowfall, Cairngorms National Park, Scotland. February
Capercaillie (Tetrao urogallus) male displaying in snow, Cairngorms NP, Scotland, February 2009
Capercaillie (Tetrao urogallus) male, head portrait, Strathspey, Cairngorms NP, Scotland, January 2009
Red deer (Cervus elaphus) stag under trees calling during rut in morning, Klampenborg Dyrehaven, Denmark, September 2008
Red deer (Cervus elaphus) stag calling during rut, light mist at sunrise, Klampenborg Dyrehaven, Denmark, September 2008. WWE INDOOR EXHIBITION
Red deer (Cervus elaphus) calling at sunset, during rut, Klampenborg Dyrehaven, Denmark, September 2008. WWE INDOOR EXHIBITION
Male Great bustard (Otis tarda) displaying, La Serena, Extremadura, Spain, April 2009
RF- Male Great bustard (Otis tarda) displaying, La Serena, Extremadura, Spain. April
Red deer (Cervus elaphus) stag calling during rut, Klampenborg Dyrehaven, Denmark, September 2008. WWE BOOK. WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS *PRESS IMAGE
Male Capercaillie (Tetrao urogallus) displaying in front of camera in a wintry pine forest, Cairngorms National Park, Scotland, February 2009 WWE BOOK, WWE OUTDOOR EXHIBITION
Gannet (Morus bassanus) courtship, Saltee Islands, County Wexford, Ireland, June 2009
Gannets (Morus bassanus) pair, Saltee Islands, County Wexford, Ireland, June 2009
European green toad (Bufo viridis) calling, vocal sac inflated, with eyes closed, in a pond (at 2, 711m) Adylsu valley, side valley to Baksan valley and Elbrus, Caucasus, Russia, June 2008
European Roller (Coracias garrulus) calling, Pusztaszer, Hungary, May 2008
Great skua (Stercorarius skua) displaying, Shetland Isles, Scotland, UK, July
Black grouse (Tetrao tetrix) male in flight at lek, Cairngorms NP, Grampian, Scotland, UK, April
Close up of Capercaillie (Tetrao urogallus) male displaying in pine forest, Cairngorms NP, Highlands, Scotland, UK, February
Black grouse (Tetrao tetrix) male displaying at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (tetrao tetrix) males fighting at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (Tetrao tetrix) males displaying at a lek site at dawn, Cairngorms National Park, Scotland, UK, April
Black necked grebe (Podiceps nigricollis) pair at night, performing their courtship dance during the mating season, with street lights reflected in the water. The Netherlands.April 2014
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April
Common / Eurasian crane (Grus grus) displaying. Lake Hornborga, Sweden, April
RF- Black Grouse (Lyrurus tetrix) displaying. Finland. March
Common cranes (Grus grus) displaying, Hula Valley, Northern Israel, January
Adult male Capercaille (Tetrao urogallus), displaying in the snow, Scotland, December 2010
Male Little bustard (Tetrax tetrax) displaying, Catalonia, Spain, May
RF- Puffins (Fratercula) displaying. Skomer Island, Wales, UK. April
