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Walia ibex (Capra walie) herd, Simien Mountains National Park, Ethiopia
Great Frigatebird (Fregata minor) in flight over Midway Atoll. Midway
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Ice fragment on crack in Lake Baikal ice, Siberia, Russia, March
Roseate Spoonbills (Platalea ajaja) in flight, Pantanal, Mato Grosso do Sul, Brazil
Hyacinth Macaw (Anodorhynchus hyacinthinus) small group in flight, Brazil
Flock of Stellers eiders (Polysticta stelleri) in flight, Batsfjord, Norway, March
Commen raven (Corvus corax) flying in falling snow, Vardo, Norway, March
Sanderling (Calidris alba) running along the tide-line, reflected in the mud, Norfolk, England, UK. December
Yellow lipped sea snake (Laticauda colubrina) dives back down to the reef after surfacing for breath of air. Apo Island, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
RF - Long exposure of a school of yellowtail surgeonfish (Prionurus punctatus), these fish school to help them access the best feeding areas, which are controlled by aggressive damselfish
RF- Two Greater flamingos (Phoenicopterus ruber) flying across sunset sky, Namibia
Two Great egrets (Ardea alba) wading, silhouetted at dawn, Keoladeo National Park, Bharatpur, Rajasthan, India
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at sunrise. Kasanka National Park, Zambia
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Blacktip reef sharks (Carcharhinus melanopterus) swimming in shallow crystal clear water, Aldabra Atoll, Seychelles, Indian Ocean
Wild yak (Bos mutus) running, Kekexili, Qinghai, Tibetan plateau, China, December
European rabbits (Oryctolagus cuniculus) juveniles emerging from burrow, Cheshire, UK May
A flock of Greylag Goose (Anser anser) flying in formation above misty fields. The Netherlands, March
Mustang / wild horses, second year cremello colt Claro play fighting with bay, McCullough Peak herd, Wyoming, USA, July 2009
Western grebe (Aechmophorus occidentalis) pair swimming, with reflections, Bolsa Chica Ecological Reserve, California, USA, February
Little Auks (Alle alle) flying low above surface in front of iceberg. Spitsbergen, Svalbard, Norway, July 2010. Highly commended in the Birds category in the GDT competition 2011
Emperor penguins (Aptenodytes forsteri) males dive through openings in the sea ice to take their first swim in over 4 months as they have spent the bleak winter incubating their eggs
Green turtle (Chelonia mydas) from underneath swimming through the surface, Mayotte Island, Comores, Indian Ocean
RF- African elephant family (Loxodonta africana) crossing desert landscape. Namibia, Kaokoland. Endangered species
Tiger (Panthera tigris tigris), male portrait, Bandhavgarh, India, February 2013
Common dolphin (Delphinus delphis) surfacing, Atlantic ocean, Portugal, September
A shoal of Golden grey mullet (Liza aurata) swim in the surf breaking on the shore of El Hierro Island. Canary Islands, Spain. East Atlantic Ocean
RF - Maldive anemonefish (Amphiprion nigripes) swimming above their host Magnificent sea anemone (Heteractis magnifica) on coral reef
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
RF- Arabian Oryx (Oryx leucoryx) Dubai Desert Conservation Reserve, Dubai, UAE
RF- Eastern white pelicans (Pelecanus onocrotalus) taking off from water, Danube delta rewilding area, Romania. May
RF- Great grey owl (Strix nebulosa) in flight. Kuhmo, Finland
Lyrtetail anthias (Pseudanthias squamipinnis) in water column with Peach anthias (Pseudanthias dispar) in background. Indonesia, October
Polar bear (Ursus maritimus) walking along coast of Wrangel Island, Far Eastern Russia, October
RF - Bannerfish (Heniochus diphreutes) schooling in coral reef. Long exposure. North Male Atoll, Maldives. Indian Ocean
Demoiselle crane (Grus virgo or Anthropoides virgo), large flock at their wintering site, Rajasthan, India
Great egret (Ardea alba) in flight, Florida, USA, March
RF- White stork (Ciconia ciconia) in flight, La Serena, Extremadura, Spain. March
RF- White-tailed Sea Eagle (Haliaeetus albicilla) turning in flight, Flatanger, Norway, August
RF- White tailed sea eagle (Haliaeetus albicilla) in flight, Portree, Skye, Inner Hebrides, Scotland, UK. June
Pine marten (Martes martes) leaping from branch, orange glow in sky behind from the lights of Inverness. Black Isle, Scotland, UK, November. Photographed by camera trap
Male black rhinoceros (Diceros bicornis) silhouetted against clouds, Masai-Mara Game Reserve, Kenya. March
Ptarmigan (Lagopus mutus) in winter plumage in flight, Utsjoki, Finland, April
Red-crowned Crane (Grus japonensis) in flight, Hokkaido, Japan, February
Greater flamingos (Phoenicopterus ruber) in flight over sand dune, Donana National Park, Andalusia, Spain, March
