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Chilean flamingos (Phoenicopterus chilensis) in flight over mountain peaks with glacier in the distance, Torres Del Paine National Park, Chile
Rose-ringed / ring-necked parakeets (Psittacula krameri) in flight on way to roost in an urban cemetery, London, UK. January
Female Harpy Eagle (Harpia harpyja) in flight. Returning to its nest. Pousada Currupira d Araras, south west Brazil
Curlews (Numenius arquata) group flying over the sea during storm. Wales, UK December. Highly commended, Habitat division, British Wildlife Photography Awards (BWPA) competition 2012
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Black-throated Diver (Gavia arctica) Rokua, Finland, May
Line of Leaf-cutter ants (Atta sp) carrying leaves, digital composite
Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Whale Shark (Rhincodon typus) passing by baitball, Santa Maria Island, Azores, Portugal, September 2008 Exclusive Japanese calendar rights for 2014
RF- African elephant (Loxodonta africana) at sunset. Chobe National Park, Botswana. Endangered species
Adelie penguin (Pygoscelis adeliae) diving near ice flow, Antarctica. Small reproduction only
RF- Wild male Jaguar (Panthera onca palustris) running through shallows of backwater of Cuiaba River in late afternoon sunlight. Northern Pantanal, Brazil
Blue-lined snapper (Lutjanus kasmira) shoal. Cocos Island National Park, Costa Rica
Large shoal of Horse mackerel (Megalaspis cordyla) schooling. Rinca, Komodo National Park, Indonesia
Shag (Phalacrocorax aristotelis) in flight over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Canada goose (Branta canadensis) adult swimming leading five goslings, Ithaca, New York, USA
Juvenile male Peregrine falcon (Falco peregrinus) in flight chasing his parent who has Feral pigeon (Columba livia) kill in claws, Bristol, England, UK, June
RF- Small group of Pink-footed geese (Anser brachyrhynchus) in flight at dawn, The Wash Estuary, Norfolk, England, UK, October 2011
Chilean flamingos (Phoenicopterus chilensis) group of three in flight, Lago Amarga, Torres del Paine National Park, Patagonia, Chile
Female Capybara (Hydrochoerus hydrochaeris) swimming in line with young after escaping a Jaguar attack (Panthera onca palustris) in a lagoon off the Paraguay River
Giant anteater (Myrmecophaga tridactyla) walking in habitat, Hato El Cedral. Llanos, Venezuela
Tibetan Wild Ass / Kiang, (Equus kiang) running, ChangThang, Tso Kar lake, at altitude of 4600m, Ladakh, India
Chamois (Rupicapra rupicapra) running down ridge, in early morning light. Lausons Valley, Gran Paradiso National Park, Italy, September
Golden Eagle (Aquila chrysaetos) in flight rear view, Kuusamo, Finland, February
Steppe eagle (Aquila nipalensis) in flight, Oman, November
Golden Eagle (Aquila chrysaetos) in flight, snowing, Utajarvi, Finland, February
Mountain hare (Lepus timidus) running in half summer coat, Peak District, UK April
School of large Atlantic bluefin tuna (Thunnus thynnus) captive in growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Lesser flamingo flock (Phoeniconaias) flying over lake with green algae, aerial view, Rift valley, Kenya
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Blue shark (Prionace glauca) Pico Island, Azores, Portugal, Atlantic Ocean, July
Dall sheep (Ovis dalli) yearling lamb running across alpine tundra, Denali National Park, Alaska, USA
Great egret (Ardea alba) flying across lake, Ranthambhore National Park, India
RF - Red-and-Green Macaws or Green-winged Macaws (Ara chloropterus) in flight over forest canopy. Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Two Masai Giraffes (Giraffa camelopardalis tippelskirchii) heads silhouetted against sky, above tree line at dusk, Serengeti NP, Tanzania, East Africa, January
Red-and-green macaw (Ara chloropterus) in flight, Pantanal, Brazil. August
Male Red winged blackbird (Agelaius phoeniceus) taking off, New York, USA, May
Big-eye fish (Priacanthus hamrur). Egypt, Red Sea
Coral reef scenery with Gorgonian and shoal of Pygmy / Glassy sweepers (Parapriacanthus guentheri) Egypt, Red Sea
King penguin (Aptenodytes patagonicus) colony. Right Whale Bay, South Georgia. September
RF- Male Emperor Dragonfly (Anax imperator) in flight, Arne RSPB reserve, Dorset, England, UK, July. Did you know? Dragonflies spend up to six years as drab aquatic nymphs for only a few weeks as
White-tailed Eagle (Haliaeetus albicilla) in flight with mountains in background, Hokkaido, Japan, February
Whale shark (Rhincodon typus) with shoal of anchovies staying near to avoid predation. Los Roques, Venezuela. Caribbean
Musk ox (Ovibos moschatus) two covered in snow, Wrangel Island, Far Eastern Russia, September
Sockeye Salmon (Oncorhynchus nerka) travelling upstream on spawning migration, Adams river, British Columbia, Canada, October Taken for the Freshwater Project
School of Bluespine unicornfish (Naso unicornis) gathered during the summer breeding season. Yolanda Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
School of young European perch (Perca fluviatilis) in Altausseer lake, Austria, July
