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Remarkable Rocks at night, Flinders Chase National Park, Kangaroo Island, South Australia State, Australia, September 2011
Sand dunes on Agilos Kopa, Nagliai Nature Reserve, Curonian Spit, Lithuania, June 2009. WWE BOOK. WWE INDOOR EXHIBITION
Iceberg in the sea outside the Kangia Ilulissat Icefjord. Iceberg from the Sermeg Kujalleg Glacier, UNESCO World Heritage Site, Kalaallit Nunaat, Greenland, August 2014
Greenland shark (Somniosus microcephalus) with parasitic copepod (Ommatokoita elongata), under ice, Lancaster Sound, Nunavut, northern Baffin Island, Canada, Arctic
Coalition of male Lions (Panthera leo) in grassland before storm, Masai-Mara game reserve, Kenya
Golf-ball sized Painted frogfish (Antennarius pictus) waiting to ambush prey disguised as an orange sponge. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Japanese fireflies (Luciola cruciata) in flight at night, endemic species, Yaku-shima UNESCO World Heritage Site, Kagoshima, Japan, May
Detail of melting ice on Jokulsarlon beach. Southern Iceland, Europe, November 2012
Silhouette of weathered, windswept hawthorn tree on stormy morning sky, Near Lynton, Devon, UK. October 2008
Scenic view of a bull kelp forest (Nereocystis luetkeana) with sunlight shining through the fronds. Browning Pass, Port Hardy, Vancouver Island, British Columbia. Canada. North East Pacific Ocean
Mountain Hare (Lepus timidus) on moorland with white winter coat, Kinder Scout, Peak District National Park, Derbyshire, UK, February
Deer mouse (Peromyscus maniculatus) on giant puffball mushroom, watching mosquito in the moonlight. Blackfoot Valley, Western Montana, USA, July
Tawny nurse shark (Nebrius ferrugineus) Vaavu Atoll, Maldives, Indian Ocean
Leatherback Turtle (Dermochelys coriacea) returning to the sea after egg laying. Cayenne, French Guiana, June
Group of lions (Panthera leo) at night, Masai Mara, Kenya. Photographed using Starlight Camera technology without artificial lighting
Moai stone statue at dusk on the slopes of the quarry at Rano Raraku volcano, Easter Island Pacific ocean, November 2004
Peregrine falcon (Falco peregrinus) female, flying towards nest box with three chicks, aged 18 days, inside, Sagrada Familia Basilica, Catalonia, Spain. May
Strinati's cave salamander (Speleomantes strinatii) inside a cave with the cave's entrance visible on the right of the image. Genoa, Italy
Praying mantis (Mantidae) with water vapour from cloud. Taken at high altitude hill station, Himalayas, India
Juvenile Puma (Puma concolor) captured on a camera trap in the Ecuadorian Choco rainforest, one of the most biodiverse and threatened rainforests on Earth. Mashpi Reserve, Pichincha, Ecuador, April
Female leopard (Panthera pardus) drinking at night, Zimanga private game reserve, KwaZulu-Natal, South Africa
RF - Schoolmaster snapper (Lutjanus apodus) swimming through underwater caverns within a coral reef, Grand Cayman, Cayman Islands, Caribbean Sea
Red fox (Vulpes vulpes) walking through Highgate Cemetery Mausoleum at night, London, UK. August
Red fox (Vulpes vulpes) vixen looking in through window of traditional cottage house, Vertes Mountains, Hungary
Female Red fox (Vulpes vulpes) cautiously staring at the camera from the base of a moss-covered tree, Hungary. January
Caribbean reef squid (Sepioteuthis sepioidea) portrait reflecting off the surface of the ocean at night, Eleuthera, Bahamas
European river otters (Lutra lutra) reflection in the water at night. Lincolnshire, UK, September
European river otter (Lutra lutra) in mist reflected in water at night. Lincolnshire, UK, September
Mountain peaks and ridges in evening light, stormy skies above. Zanskar Mountains, Ladakh, India. September 2011
Little owl (Athene noctua) taking shelter from the rain, Arcos de la Frontera, southern Spain. April
Riisitunturi in winter with Aurora Borealis, Kuusamo, Lapland, Finland. February
Dry stone wall, near Birch tor, Dartmoor NP, Devon. September 2008
Hut circle remains and large circular granite slab, against stormy sky, Merrivale, Dartmoor NP, Devon, UK. September 2008
Badger (Meles meles) crossing a burn in alder woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, August
Leopard (Panthera pardus) walking along waterhole, reflected in the water at dusk. Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Solar eclipse, Finland, 20 March 2015
Ring tailed lemur (Lemur catta) backlit in late afternoon light, Berenty Private Reserve, southern Madagascar
Head portrait of grey Andalusian stallion in dark barn, Northern France, Europe
Perito Moreno Glacier, Patagonia, Southern Argentina. January 2014
Nile crocodile (Crocodylus niloticus) at night, Zimanga private game reserve, KwaZulu-Natal, South Africa
Anhumas Abyss, a 72 meters deep cave, with underwater limestone cones up to 20 meters high, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil November 2016
Reindeer (Rangifer tarandus) reindeer silhouetted against the skyline, reintroduced Cairngorm Reindeer Herd, Cairngorm National Park, Speyside, Scotland, July
Malham Moor, with stormy skies over Winskill and Ribblesdale with Pen-y-Ghent shrouded in cloud, Yorkshire Dales National Park, UK. September 2017
Ganesh Himal at sunrise, under dark sky. Seen from Cholangpati, Cholangpati, Langtang - Gosaikund region, Nepal, November 2009
Smoke from bush fire in Pilbara region, Western Australia. August 2009
Cow parsely (Anthriscus sylvetris) in damp meadow, Lithuania, June 2009
Cuillins Mountains seen from Elgol, Isle of Skye, Scotland, UK, November 2011