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White tent making bat (Ectophylla alba) roosting in tree, La Selva Field Station, Costa Rica
Great green macaw (Ara ambiguus) flying, La Selva Field Station, Costa Rica
Travellers palm (Ravenala madagascariensis) clos up, Costa Rica
Canon ball tree fruit (Couroupita guianensis) Costa Rica
Snowcap hummingbird (Microchera albocoronata) visiting Porterweed (Stachytarpheta sp), Braulio Carrillo National Park, Talamancan montane forest, Costa Rica. Endemic species
Strawberry poison frog (Oophaga pumilio) pair mating, Sarapiqui, Heredia, Costa Rica
Common / Green iguana (Iguana iguana) portrait, Sarapiqui, Heredia, Costa Rica
Chocoan parrot snake (Leptophis bocourti) on heliconia flower with fly in background, Sarapiqui, Heredia, Costa Rica
Black-headed bushmaster (Lachesis melanocephala) with tongue extended, Siquirres, Limon, Costa Rica
Smooth helmeted iguana (Corytophanes cristatus) portrait, Sarapiqui, Heredia, Costa Rica
Misfit / Jumping leaf frog (Agalychnis saltator) looking out from behind leaf, Siquirres, Limon, Costa Rica
Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica
Fer-de-lance (Bothrops asper) camouflaged on the rainforest floor. Corcovado National Park, Osa Peninsula, Costa Rica
Fleischmanns glassfrog (Hyalinobatrachium fleischmanni) male attending eggs on the underside of a leaf overhanging a rainforest stream
Jumping spider (Salticidae) hunting among vegetation, San Jose, Costa Rica
Silk moth (Automeris zugana) sequence 2 of 2, with wings open to reveal eyespots, a means of deterring predators, photographed on a white background in mobile field studio
Red-eyed leaf frog (Agalychnis callidryas) male, Central Caribbean foothills, Costa Rica
Granular poison frog (Oophaga granulifera) Osa Peninsula, Costa Rica. Vulnerable IUCN Red List species
Lemur leaf frog (Agalychnis lemur) at night, Central Caribbean foothills, Costa Rica. IUCN Red List critically endangered species
Hourglass treefrog (Dendropsophus ebraccatus) male with inflated vocal sac calling at night, Central Caribbean foothills, Costa Rica
Leafcutter ants (Atta sp) colony harvesting a banana leaf, Costa Rica. 3rd place in the Insects and Spiders category of the Terre Sauvage Nature Images Awards competitions 2015
Great Green Macaw (Ara ambiguus) close up of feathers, El Manantial Macaw Sanctuary, Costa Rica. Captive
Morpho butterfly (Morpho peleides) photographed on white background. Costa Rica, March
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Scarlet macaw (Ara macao) flying, blurred motion. Costa Rica
Hoffmanns Two-toed Sloth (Choloepus hoffmanni) orphaned babies in rehabilitation program, Aviarios Sloth Sanctuary, Costa Rica
Leaf cutter ant (Atta sp) female worker carrying leaf to nest, with a smaller minor ant riding. The minors job is to protect her sister from parasitic flies. Costa Rica, December
Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December
Green crowned brilliant hummingbird (Heliodoxa jacula) male taking off to feed at feeder in garden, another waiting perched on lichen covered branch, Costa Rica, January
Three Brown-hooded Parrots (Pyrilia haematotis) one taking off from branch, in the rain Laguna del Lagarto, Santa Rita, Costa Rica
Violet Sabrewing Hummingbird (Campylopterus hemileucurus) male hovering at feeder, Bosque de Paz, Costa Rica, Central America
Highland cloud forest in the Talamanca Mountains of Costa Rica, April 2015
RF - Granular poison frog (Oophaga granulifera). Osa Peninsula, Costa Rica. Vulnerable species
RF - Lowland rainforest, Osa Peninsula, Costa Rica
Cup fungus (Cookeina sp) growing on decaying wood on the rainforest floor, Corcovado National Park, Osa Peninsula, Costa Rica
Aerial view of Delta Sierpe River Terraba, Corcovado National Park, Osa Peninsula, Puntarenas Province, Costa Rica. December 2014
Baby Olive Ridley sea turtle (Lepidochelys olivacea) struggles against the swell to swim away from the beach where it hatched, Ostional, Costa Rica. October
Granular poison frog (Oophaga granulifera) resting on leaf at night, Uvita, Costa Rica
Rainforest toad (Incilius aucoinae) male, portrait, Osa Peninsula, Costa Rica
Jumping spider (Lyssomanes sp. ) female, guarding its eggs on a leaf, Osa Peninsula, Costa Rica. Focus stackedJumping spider (Lyssomanes sp.) female, guarding its eggs on a leaf, Osa Peninsula, Costa Rica. Focus stacked
Sylvia's tree frog (Cruziohyla sylviae), Veragua rainforest, Costa Rica
Owl Butterfly (Caligo memnon) caterpillar heads, Costa Rica
Adult King Vulture (Sarcoramphus papa) feeding at a carcass. Laguna del Lagarto, Boca Tapada, Caribbean slope, Costa Rica, Central America
Male Resplendent Quetzal (Pharomachrus mocinno) in cloud forest. Los Quetzales National Park, Savegre River Valley, Talamanca Range, Costa Rica, Central America
Male Violet Sabrewing (Campylopterus hemileucurus) hovering / in flight sequence. Montane forest, Bosque de Paz, Caribbean slope, Costa Rica, Central America
Pair of Great curassow (Crax rubra), La Selva Biological Station, Costa Rica
Ocelot (Leopardus pardalis) sitting on forest floor, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Ocelot (Leopardus pardalis) hiding amongst tree roots, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats