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Ghost glass frog (Centrolenella ilex) portrait, Costa Rica
Leaf cutter ants (Atta sp) carrying colourful plant matter, reflected in water, Laguna del Lagarto, Santa Rita, Costa Rica
Leaf cutter ants (Atta sp) carrying plant matter, Costa Rica
Splendid leaf frog (Cruziohyla calcarifer) La Selva Field Station, Costa Rica
RF - Keel-billed toucan (Ramphastos sulfuratus) perched on mossy branch. Boca Tapada, Costa Rica. (This image may be licensed either as rights managed or royalty free.)
Scarlet macaws (Ara macao) La Selva, Costa Rica
Nicaraguan giant glass frog (Espadarana prosoblepon) La Selva Field Station, Costa Rica
Black-crowned Central American squirrel monkey (Saimiri oerstedii) pair playing with each other, one with hand over eyes of other, Manuel Antonio National Park, Quepos, Costa Rica
Black-crowned Central American squirrel monkey (Saimiri oerstedii) pair close-up Manuel Antonio National Park, Quepos, Costa Rica
Resplendent quetzal (Pharomachrus mocinno) male, Talamanca mountains, Costa Rica
Violet sabrewing hummingbird (Campylopterus hemileucurus) male hummingbirds territorial fight at dawn, Turrialba, Costa Rica
Reticulated glass frogs (Hyalinobatrachium valerioi) pair in amplexus, Osa Peninsula, Costa Rica
Snake-mimic caterpillar (Hemeroplanes triptolemus) a hawkmoth caterpillar that resembles a viper when threatened, on branch, La Selva Biological Station, Costa Rica
Male Mantled howler monkey (Alouatta palliata) on foot bridge, La Selva Biological Station, Costa Rica
Ocelot (Leopardus pardalis) climbing a tree trunk Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Keel-billed toucan (Ramphastos sulfuratus) feeding, tossing fruit seed in beak, perched on branch. Boca Tapada, Costa Rica. Sequence 2/2
White-faced Capuchin (Cebus capucinus imitator) Carara National Park, Costa Rica
Scarlet Macaw (Ara macao) in flight over the rainforest Corcovado National Park, Osa peninsula, Costa Rica
Bairds tapir (Tapirus bairdii) walking along a beach in Corcovado National Park, Costa Rica. Endangered
View of the Rio Celeste waterfall, tropical rainforest of Tenorio Volcano National Park, Costa Rica
White-faced capuchin monkey (Cebus capucinus) in Tenorio Volcano National Park, Costa Rica
Eyelash pit viper (Bothriechis schlegelii) (Viperidae: Crotalinae). Distictive yellow / orange oropel form. Arboreal species resting on wild ginger flower in mid-altitude rainforest under storey
Bigeye trevally (Caranx sexfasciatus), a large shoal. Cocos Island National Park, Costa Rica
Rocks and coastal rainforest. Cocos Island National Park, Costa Rica. 2018
Squirrelfish (Holocentrus sp) shoal amongst rocks. Near Cocos Island National Park, Costa Rica
Blue-lined snapper (Lutjanus kasmira) shoal. Cocos Island National Park, Costa Rica
Fish seeking shelter around floating tree in open ocean. Off coast of Cocos Island National Park, Costa Rica
Silky shark (Carcharhinus falciformis). Cocos Island National Park, Costa Rica
Rufous motmot (Baryphthengus martii) perched on branch. Rancho Naturalista, Cartago, Costa Rica
Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica
Rivolis hummingbird (Eugenes fulgens) nectaring on Fuchsia (Fuchsia sp) flower. Costa Rica
Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica
Violet sabrewing hummingbird (Campylopterus hemileucurus) nectaring on Orchid. Costa Rica
Red-legged honeycreeper (Cyanerpes cyaneus) two males on barbed wire. Costa Rica
White-throated magpie-jay (Calocitta formosa) Nicoya peninsula, Costa Rica. February
Adult Eyelash Pit Viper or Eyelash Viper (Bothriechis schlegelii) (Viperidae: Crotalinae). Distictive yellow / orange oropel form
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Bairds tapir (Tapirus bairdii) Corcovado National Park, Costa Rica, May. Endangered
Hoffmanns Two-toed sloth (Choloepus hoffmanni) mother and baby, aged 2 months, in tree, Costa Rica. Rescued and released by Aviarios Sloth Sanctuary
Satiny Parrot snake (Leptophis depressirostris) Santa Rita, Costa Rica
Fungal mycelium from an unidentified species of fungus, growing across a decomposing leaf. The mycelium is the vegetative part of the fungus and is made up of a mass of thread-like hyphae
Moss mimic stick insect (Trychopeplus laciniatus) showing amazing camouflage on mossy vine, photographed in tropical rainforest in mobile field studio against a white background
Strawberry poison frog (Oophaga pumilio) photographed on a white background in mobile field studio, Central Caribbean foothills, Costa Rica
Ghost glass frog (Sachatamia ilex), Costa Rica
Wild Buffons macaw x Scarlet macaw (Ara ambiguus x Ara macao) hybrid, Costa Rica
Pair of Great curassow (Crax rubra), La Selva Biological Station, Costa Rica
Ocelot (Leopardus pardalis) in rainforest, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Ocelot (Leopardus pardalis) looking up, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Ocelot (Leopardus pardalis) high up in tree, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Rufous-tailed hummingbird (Amazilia tzacatl) feeding from Heliconia flower (Heliconia champneiana) La Selva Field Station, Costa Rica
Northern tamandua (Tamandua mexicana) sleeping in a tree Corcovado National Park, Osa Peninsula, Puntarenas Province, Costa Rica
Scarlet Macaw (Ara macao) couple fighting in a tree Corcovado National Park, Osa peninsula, Costa Rica
Bairds tapir (Tapirus bairdii) browsing on leaves, rainforest, Corcovado National Park, Costa Rica. Endangered
Central American spider monkey (Ateles geoffroyi) juvenile, Corcovado National Park, Costa Rica. January
Bairds tapir female (Tapirus bairdii) with calf, rainforest, Corcovado National Park, Costa Rica. Endangered
Eyelash Viper (Bothriechis schlegelii) Distictive yellow / orange oropel form. Arboreal species resting in mid-altitude rainforest under storey. Caribbean slope, Costa Rica, Central America
Adult King Vulture (Sarcoramphus papa) feeding at a carcass. Laguna del Lagarto, Boca Tapada, Caribbean slope, Costa Rica, Central America
RF - Male Violet Sabrewing hummingbirds (Campylopterus hemileucurus) hovering in flight sequence. Montane forest, Bosque de Paz, Caribbean slope, Costa Rica
Purple throated mountain gem hummingbird (Lampornis calolaemus) hovering as it nectars on Porterweed (Stachytarpheta frantzii). Costa Rica
Red-legged bumble-moth (Homoeocera gigantea) on Lichen covered tree trunk. Costa Rica
Cup fungus (Cookeina sp) growing on decaying wood on the rainforest floor, Corcovado National Park, Osa Peninsula, Costa Rica
Leaf cutter ants (Atta sp) carrying piece of leaf, Costa Rica
White tent making bat (Ectophylla alba) roosting in Heleconia leaf, La Selva Field Station, Costa Rica
Resplendent quetzal (Pharomachrus mocinno) male perched on branch. Costa Rica
Leaf-nosed bat (Phyllostomidae sp) flying towards Banana (Musa sp) flower to feed. Costa Rica
Green-crowned brilliant (Heliodoxa jacula), three feeding on Lobster-claw (Heliconia sp) flower in rain. Costa Rica
Montezuma oropendola (Psarocolius montezuma) hanging upside down as part of a courtship display. Costa Rica
Double-crested basilisk (Basiliscus plumifrons) running across water surface, Santa Rita, Costa Rica
Leaf mimic bush cricket or katydid (unknown species, family Tettigoniidae) in the middle of moulting its skin. Moulting from one sub-adult stage (instar) to another is known as ecdysis
White fronted capuchin monkey (Cebus capucinus) feeding, March, Costa Rica
Puma (Puma concolor) looking upwards, Corcovado National Park, Costa Rica, May
Adult King Vulture (Sarcoramphus papa) Laguna del Lagarto, Boca Tapada, Caribbean slope, Costa Rica, Central America
Female Red-eyed Tree Frog (Agalychnis callidryas) - Caribbean slope race (blue flanks) - obscured in rainforest foliage. Mid-altitude rainforest near Aranal, central Costa Rica
White-headed capuchin (Cebus capucinus) walking on giant palm-tree, Costa Rica
Masked tree frog (Smilisca phaeota) El Arenal region, Costa Rica
Proboscis bats (Rhynchonycteris naso) day roosting, Costa Rica, March 2014
Leaf-cutter ants (Atta cephalotes) carrying pieces of leaf that they have harvested back to their underground fungus garden in their nest, Osa Peninsula, Costa Rica
Slime mould (Physarum sp) plasmodium growing across rotting wood, Osa Peninsula, Costa Rica
Spiny predatory katydid (Listroscelis sp) photographed on a white background in mobile field studio, Osa Peninsula, Costa Rica
Clearwing moth (Cosmosoma teuthras) photographed on a white background in mobile field studio, Cordillera de Talamanca mountain range, Caribbean Slopes, Costa Rica
Army ants (Eciton sp. ) Costa Rica. February 2015Army ants (Eciton sp.) Costa Rica. February 2015
Orange-barred sulphur butterfly (Phoebis philea) pe. Costa Rica
Great curassow (Crax rubra), Costa Rica