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Slate pencil sea urchin (Heterocentrotus mammillatus) on a reef with Black triggerfish (Melichthys niger) and a Bluestripe snapper (Lutjanus kasmira) swimming above, Maui, Hawaii, Pacific Ocean
Bluespotted ribbontail ray (Taeniura lymma) searching for food over the coral reef at night, Triton Bay, West Papua, Indonesia, Pacific Ocean
Cleaning shrimp (Periclimenes sp) on a coral reef, close up, Triton Bay, West Papua, Indonesia, Pacific Ocean
Soft coral (Dendronepthya sp) garden, Iris Strait, Triton Bay, Indonesia, Pacific Ocean
School of Yellow-ribbon sweetlips (Plectorhinchus polytaenia) swimming with the current, Raja Ampat, Indonesia, Pacific Ocean
Hawksbill turtle (Eretmochelys imbricata) cloze up on coral reef, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Blackspotted puffer (Arothron nigropunctatus) curled up on the reef for the night, with a transparent cleaner shrimp (Urocaridella antonbrunii) inspecting its tail, Philippines
Female California sheephead (Semicossyphus pulcher) and miscellaneous reef fish swimming in a forest of Giant kelp (Macrocystis pyrifera) off Santa Barbara Island, California, USA
A Green sea turtle (Chelonia mydas) at surface to breathe, over shallow reef, Hawaii USA
Pair of mating Big blue octopuses (Octopus cyanea) where the male, on the right, is fertilising the female using its fertilising tentacle. Indian ocean, Maldives
A young lionfish (Pterois volitans) swimming near the surface (hunting silversides, not pictured) close to dusk. Beacon Rock, Sha ab Mahmood, Sinai, Egypt. Red Sea
Female Yellowback anthias (Pseudanthias evansi), Yap, Micronesia
RF - Portrait of Spinecheek anemonefish (Premnas biaculeatus) in its host Bubble-tip anemone (Entacmaea quadricolor) on a coral reef. Raja Ampat, West Papua, Indonesia. Ceram Sea
Portrait of a Secretary blenny (Acanthemblemaria maria), living in a hole in the base of a purple common sea fan, on a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies
Colourful soft corals, Dead mans fingers (Alcyonium digitatum) and common sea urchins (Echinus esculentus) mass beneath a kelp forest (Laminaria hyperborea)
Tiny (1cm) Pygmy seahorse (Hippocampus bargibanti) hides in the branches of its home seafan (Muricella sp)Tiny (1cm) Pygmy seahorse (Hippocampus bargibanti) hides in the branches of its home seafan (Muricella sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea
False clown anemonefish (Amphiprion ocellaris) looking out of a magnificent sea anemone (Heteractis magnifica) on a coral reef. Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Yellow boxfish (Ostracion cubicus) adult, Yap, Micronesia
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Common octopus (Octopus vulgaris) hunting on a reef off Eleuthera Island, Bahamas
Split level image of Leather corals (Sarcophyton sp. ) flourishing in shallow water closeSplit level image of Leather corals (Sarcophyton sp.) flourishing in shallow water close to uninhabited coral islands. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea
Caribbean reef shark (Carcharhinus perezi) patrolling coral reef with Schoolmaster snapper (Lutjanus apodus) and other fish. Caribbean Sea off Gardens of the Queen National Park, Cuba
Schoolmaster snapper (Lutjanus apodus) school in the reef, Little Cayman island, Cayman Islands, Caribbean
Table coral reef in shallow waters, Gaafu Alifu Atoll, Maldives, Indian Ocean
Great hermit crab (Dardanus arrosor). Santa Maria Island, Azores, Portugal, Atlantic Ocean
Bridled anemonefish (Amphiprion frenatus) in a Bulb-tenacle sea anemone (Entacmaea quadricolor) in reef of finger coral, Yap, Micronesia
Colourful Soft corals (Dendronephthya hemprichi) with reef fishes, including Golden damselfish (Amblyglyphidodon aureus) at Hin Daeng
Coral reef scene, with Philippines chromis (Chromis scotochiloptera), hard coral (Acropora sp.) and purple elephant ear sponges (Ianthella basta)
Coral grouper (Cephalopholis miniata) waits in ambush, hiding against a seafan (Annella sp.) hunting silversides (Atherinidae), with schooling bigeye snapper (Lutjanus lutjanus) behind
RF - Maldive anemonefish (Amphiprion nigripes) swimming above their host Magnificent sea anemone (Heteractis magnifica) on coral reef
Mexican Hogfish (Bodianus diplotaenia), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Coral mushrooms, coral banks are eroded by change in sea level, tidal current and wind, Passe Femme / Femme channel, Aldabra, Indian Ocean 2006
Flame tip hermit crab (Calcinus minutus), a left-handed hermit crab that is often found in branching hard corals, Yap, Micronesia
Seafan / Red gorgonian (Paramuricea clavata) and Light-bulb sea squirt (Clavelina lepadiformis) Punta Carena, Capri Island, Sorrentine Peninsula, Italy, Tyrrhenian Sea, Mediterranean
Hermit crab (Dardanus arrosor), with Shoal of Ornate wrasse (Thalassoma pavo) Santa Maria Island, Azores, Portugal, Atlantic Ocean
Happy Eddie (Haploblepharus edwardsii) egg case / mermaids purse. False Bay, Western Cape, South Africa. June
Leopard shark (Stegostoma fasciatum) cruising along the edge of an underwater drop off, Philippines
Healthy Staghorn corals (Acropora sp. ) with various reef fishesHealthy Staghorn corals (Acropora sp.) with various reef fishes, including Moon wrasses (Thalassoma lunare), and Threespot dascyllus (Dascyllus trimaculatus)
Caribbean reef shark (Carcharhinus perezi) swimming over a coral reef, watched by a Nassau grouper (Epinephelus striatus) surrounded by common sea fans (Gorgonia ventalina)
RF - A giant trevally (Caranx ignobilis) scatters baitfish (silversides and cardinalfish) as it hunts on a coral reef. Yillet Islands, Misool, Raja Ampat, West Papua, Indonesia
Reef manta (Mobula alfredi) visiting a cleaning station on a coral reef with a school of Onespot snappers (Lutjanus monostigma)
Portrait of a Blackbar soldierfish (Myripristis jacobus) infected with Cymothoid isopod parasites. The large parasite between the eyes is a female, the smaller individuals are males
Pair of Rock beauty angelfish (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of the female during the spawning rise
View through a cavern in a coral reef, with a Small grouper (Cephalopholis cruentata) in the distance. Snapper Hole, East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Warty / Clown frogfish (Antennarius maculatus) male portrait, Anilao, Batangas, Luzon, Philippines, March
Tiny snapping shrimp (Synalpheus sp. ) looks out from within sea squirt (Didemnum molleTiny snapping shrimp (Synalpheus sp.) looks out from within sea squirt (Didemnum molle) at night on a coral reef. Piaynemo Island, Raja Ampat, West Papua, Indonesia
Caribbean reef shark (Carcharhinus perezi) swims over a coral reef, photographed through the branches of a porous sea rod (Pseudoplexaura sp.)
Orange branching sponge (Pilocaulis sp. ) growing in a coral canyon on a reef wallOrange branching sponge (Pilocaulis sp.) growing in a coral canyon on a reef wall. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea