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Slender filefish (Monacanthus tuckeri) hiding in the branches of a porous sea rod (Pseudoplexaura sp.) on a coral reef. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
RF - A threespot damselfish (Stegastes planifrons) aggressively defending its algae garden on a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea. Model Released
Zebra / leopard shark (Stegostoma fasciatum) swimming over the reef, Vaavu atoll, Maldives, Indian Ocean
Coral heads surrounded by Blacktip cardinalfish (Apogon atradorsatus)and juvenile salemas (Xenocys jessiae), Rabida Island, Galapagos
Schoolmaster snapper (Lutjanus apodus) and Bluestriped grunt (Haemulon sciurus) seeking protection in Elkhorn Coral (Acropora palmata), Jardines de la Reina / Gardens of the Queen National Park
Hawksbill turtle (Erethmochelys imbricata) feeding on tropical reef, Tulamben, North coast, Bali, Indonesia. Lesser Sunda Islands
Smallmouth grunt (Haemulon chrysargyreum) fish swim over a coral reef, Cancun, Mexico. Slow shutter speed image
Yellow stingray (Urobatis jamaicensis) on sand seabed by coral reef, The Bahamas
Broadclub cuttlefish (Sepia latimanus) over pristine coral reef, Kimbe Bay, Papua New Guinea
Pink anemonefish (Amphiprion perideraion) living in symbiotic association with Magnificent sea anemone (Heteractis magnifica), Palau
Moluccan cardinalfish (Ostorhinchus moluccensis) next to hydroids, brittlestars and sea squirts. Tulamben, Bali, Indonesia
Pair of courting Egg-eating sea snakes / Turtleheaded sea snakes (Emydocephalus annulatus) New Caledonia, Pacific Ocean
Caribbean reef shark (Carcharhinus perezi) patrolling coral reef with Schoolmaster snapper (Lutjanus apodus) and other fish. Caribbean Sea off Gardens of the Queen National Park, Cuba
Forsters hawkfish (Paracirrhites forsteri) on a coral reef. Jackson Reef, Tiran, Sinai, Egypt. Strait of Tiran, Gulf of Aqaba, Egypt
Hawksbill turtle (Eretmochelys imbricata) unusually feeding on hard coral polyps. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Green turtle (Chelonia mydas) descending after breathing at the surface. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Commensal emperor shrimp (Periclimenes imperator) moves across the colourful red patterned surface of its host Candycane sea cucumber (Thelenota rubralineata)
Split level image of Leather corals (Sarcophyton sp.) flourishing in shallow water close to uninhabited coral islands. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea
Split level image of hard coral garden flourishing in shallow water below Misool Eco Resort. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Reef manta (Mobula alfredi) female swimming close to a coral reef, while cleaner wrasse (Labroides dimidiatus), tiny by comparison, pick parasites from her lips
Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island
Tawny nurse sharks (Nebris ferrungineus) at night. Vavuu Atoll, Maldives. Indian Ocean
Midnight snapper (Macolor macularis) shelters in a cavern on a coral reef with white soft corals (Scleronephthya sp.) and solderfish. Vavuu Atoll, Maldives. Indian Ocean
Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef
Shadowfin soldierfish (Myripristis adusta) West Papua, Indonesia
Shrimpfish / Razorfish (Aeoliscus punctatus / strigatus) have adapted to a vertical posture and swim with their head downward. Lembeh, North Sulawesi, Indonesia
Shadowfin soldierfish (Myripristis adusta) schooling over coral reef. Dampier Strait, Raja Ampat, West Papua, Indonesia, March 2016
Gorgonian sea fans (Melithaea sp. ) with a large school of Pygmy sweepers (ParapriacanthusGorgonian sea fans (Melithaea sp.) with a large school of Pygmy sweepers (Parapriacanthus ransonetti) Andaman Sea, Thailand
Green mandarinfish (Synchiropus splendidus) pair courting and mating at sunset, Sulu Sea, Philippines
Kaleidoscopic image of soft coral (Dendronephthya sp.) and Lyretail anthias (Pseudanthias squamipinnis). Egypt, Red Sea
Seawhip (Cirrhipathes spiralis) Indonesia
Lyretail anthias / Goldies (Pseudanthias squamipinnis) on reef top. Egypt, Red Sea
Honeycomb / Cushion starfish (Pentaceraster alveolatus) Malapascua Island, Philippines, September
Red Sea anemonefish (Amphiprion bicinctus) in anemone. Egypt, Red Sea
Kaleidoscopic image of Coral reef scenery with gorgonian, soft corals and Lyretail anthias (Pseudanthias squamipinnis). Rinca, Komodo National Park, Indonesia
Latticetail moray (Gymnothorax buroensis) emerging from soft coral on a reef. Sangeang Island, Sumbawa, Indonesia. Flores Sea
Longnose hawkfish (Oxycirrhites typus) Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Anthias fish (Pseudanthias squamipinnis), by Fire coral (Millepora dichotoma) and soft coral, on coral reef, Shark Reef to Jolande Reef, Ras Mohammed National Park, Egypt, Red Sea
Sergeant major (Abudefduf vaigiensis), shoal on wreck of the Kingston, near Shag Rock, Strait of Gubal, Egypt, Red Sea
Split level view of shallow coral reef in the Red Sea at sunset. Gordon Reef, Sinai, Egypt. Strait of Tiran, Gulf of Aqaba, Red Sea
Vibrant Red Sea reef scene, with orange female Scalefin anthias (Pseudanthias squamipinnis) swarming in front of Fire coral (Millepora dichotoma) feeding on plankton brought to the reef by currents
A long exposure renders Bar jacks (Carangoides ruber) as ghostly trails as they swim around a sponge, on a coral reef, at night. East End, Grand Cayman, Cayman Islands
Group of Circular spadefish (Platax orbicularis) gather at a cleaning station beneath a table coral (Acropora sp.) on a coral reef. Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea
Colourful reef scene with Green tubastrea (Tubastrea micrantha), Soft corals (Scleronephthya sp)Colourful reef scene with Green tubastrea (Tubastrea micrantha), Soft corals (Scleronephthya sp.) and Scalefin anthias (Pseudanthias squamipinnis) and Magenta slender anthias (Luzonichthys waitei)
Redtail butterflyfish (Chaetodon collare) pair in front of a smooth Sea fan (Annella mollis). Butterflyfish regularly form stable pairs. Similan Islands, Thailand. Andaman Sea, Indian Ocean
Coral grouper (Cephalopholis miniata) guarding its territory on a colourful coral reef. East Of Eden, Similan Islands, Thailand. Andaman Sea, Indian Ocean
Rich coral growth, with a pair of Exquiste butterflyfish (Chaetodon austriacus), flourishes beneath the barren desert cliffs in the Red Sea. Taken at sunset. Ras Katy, Sinai, Egypt
School of Bohar snapper (Lutjanus bohar) swimming in formation along a coral reef. These fish are usually solitary and aggregate each summer in the Red Sea to spawn