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African grey hornbill (Lophoceros nasutus) calling in display, perched on tree snag. Janjanbureh, Gambia
Crab-plover (Dromas ardeola) flying with crab in beak. Oman, June
Crab-plover (Dromas ardeola) three flying, two with crabs in beaks. Oman, June
Green sea turtle (Chelonia mydas), two viewed from above. Tenerife, Canary Islands
Snakes head fritillary (Fritillaria meleagris), view into flower with linear green nectaries. In cultivation, Surrey, England, UK. April
RF - Monarch butterfly (Danaus plexippus) with life-cycle tied to the presence of milkweed plants (Asclepias sp). East Haddam, Connecticut, USA. September
Raven (Corvus corax) mobbing Gyrfalcon (Falco rusticolus), Hornoya Island, Varanger Peninsula, Norway, March
Hornet (Vespa crabro) close up of head
RF- Red squirrel (Sciurus vulgaris) portrait, in snow, Cairngorms National Park, Scotland, March 2007
RF- Red squirrel (Sciurus vulgaris) jumping. Cairngorms National Park, Scotland, March 2012
Crested drongo (Dicrurus forficatus) rear view, Berenty Reserve, Madagascar
Honey bee (Apis mellifera) in flight, Surrey, England, July
African Blue tit (Cyanistes teneriffae) Morocco, February
Female humpback whale (Megaptera novaeangliae) with its very young calf at the surface, Rurutu island, Australs Archipelago, French Polynesia. Pacific Ocean
Masai Giraffes (Giraffa camelopardalis tippelskirchi) group of three, Masai Mara Game Reserve, Kenya
Portrait of male Scalefin anthias (Pseudanthias squamipinnis) swimming in open water, to feed on plankton. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Small pearl bordered fritillary butterfly (Boloria selene) resting on grass, backlit underwing, Marsland Mouth, North Devon, UK, June
Gannet (Morus bassanus) landing in colony, Bass Rock, Scotland, UK, July
Moth (Zygaena osterodensis) feeding on flower, Viscos, Pyrenees National Park, France, July
Three Marbled white butterflies (Melanargia galathea) resting on reeds, Devon, UK, July
Silver-studded blue butterfly (Plebejus argus) pair mating, resting on a Pyrimidal orchid (Anacamptis pyramidalis) flower, Gwithian Towans, Cornwall, UK, July
tail detail of Sperm Whale (Physeter macrocephalus) Pico Island, Azores, Portugal, Atlantic Ocean
Common / Eurasian crane (Grus grus) displaying. Lake Hornborga, Sweden, April
Common / Eurasian crane (Grus grus) calling at sunset, silhouetted. Lake Hornborga, Sweden, April
Blackcap (Sylvia atricapilla) Uto Finland May
Humpback Whale (Megaptera novaeangliae) female. Tonga, South Pacific, September
Blue tits (Paris caereleus) two feeding on red berries, Uto Finland October
Wild Cat (Felis silvestris) portrait. Vosges, France, August
Bat-eared fox (Otocyon megalotis) walking, Namib-Naukluft National Park, Namib Desert, Namibia
Common darter dragonflies (Sympetrum striolatum) two females resting on reed, Dunsdon Nature Reserve, Devon, UK. August
Nudibranch (Cratena peregrina) on a sponge, Ist Island, Croatia, Adriatic Sea, Mediterranean
Song thrush (Turdus philomelos clarkei) portrait, Hungary, May
Whaleshark (Rhincodon typus) swimming and filtering fish eggs from the water. A behaviour known as ram-feeding. Isla Mujeres, Quintana Roo, Yucatan Peninsular, Mexico. Caribbean Sea
A portrait of a Mediterranean anthias (Anthias anthias) against open water. Secca Del Papa, Tavolara Marine Protected Area, Sardinia, Italy. Mediterranean Sea
Portrait of Red Sea anemonefish (Amphiprion bicinctus) against open water. Nuweiba, Sinai, Egypt
Blue shark (Prionace glauca) near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August
A portrait of a small Candy stripe shrimp (Lebbeus grandimanus) in front of its host pink anemone (out of focus because of the magnification)
RF - Bluestreak cleaner wrasse (Labroides dimidiatus) tending to Ribboned sweetlips (Plectorhinchus polytaenia). Yillet Kecil, Yillet Islands, Misool, Raja Ampat, West Papua, Indonesia
RF- Orange Tip butterfly (Anthocharis cardamines) female resting on Cuckooflower (Cardamine pratensis). North Devon, UK. April
Golden-headed lion tamarin (Leontopithecus chrysomelas) captive portrait, Apenheul Park, Netherlands
Green-veined White butterfly egg (Pieris napi) above, Orange-tip butterfly egg (Anthocharis)Green-veined White butterfly egg (Pieris napi) above, Orange-tip butterfly egg (Anthocharis cardamines) below, on Garlic mustard (Alliaria petiolata) Sussex, UK. June
Female Azure damselfly (Coenagrion puella), Vallee de la Seille (Seille Valley), Lorraine, France, June
Blue shark (Prionace glauca) as it cruises beneath the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August
Blue Tit (Paris caereleus) hanging below red berries on which it has been feeding, Uto Finland November
RF- Mexican treefrog 1+Smilisca baudinii+2 on Spanish moss. Texas, USA
Firewheel (Gaillardia pulchella) flowers at twilight, Laredo Borderlands, Texas, USA. April
