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Holkham beach at low tide, Norfolk, UK August
Footprints of an Amur / Siberian tiger (Panthera tigris altaica) in sand along the edge of the Sea of Japan, Lazovskiy Zapovednik Nature Reserve, Primorsky Krai, Russian Far East. February 1992
Palm trees line the beach at Pigeon Point, Tobago
Dramatic evening sky over Anse Source d Argent, La Digue. Seychelles. February 2007
Aerial view of the Long Wall, sand dunes along the Atlantic coast of the Namib desert, Swakopmund, Namibia, August 2008
Aerial image showing sandbanks in the Bahamas archipelago, Caribbean, February 2012
Aerial view of Everglades National Park, Florida, USA, February 2012
White horses of the Camargue, herd running through water, Camargue, Southern France
Andalusian stallion running out of the sea on beach, Ojai, California, USA
Sandstone cliffs, Rackwick Bay, Hoy, Orkney, Scotland, July 2004
Icebergs off the Antarctic Peninsula, Antarctica, February 2009, Taken on location for BBC Frozen Planet series
Gulls on the beach at sunset, The Wash, Hunstanton, Norfolk, UK, September 2011
Leatherback Turtle (Dermochelys coriacea) returning to the sea after egg laying. Cayenne, French Guiana, June
Puffins (Fratercula arctica) pair take off together to go fishing. Farne Islands, Northumberland, UK, May. Non-ex
Durdle Door on the Jurassic Coast of Dorset. England, November 2010
Aerial view of the rivers and saltmarshes of the Bahia / Bay de Cadiz Natural Park, Andalucia, Spain, March 2008
Male Panther Chameleon (Furcifer pardalis) feeding on Preying mantis in beach side vegetation. Bay of Antongil, Masoala Peninsula National Park, north east Madagascar
Marine Iguanas (Amblyrhynchus cristatus) on the south coast of Espanola Island, Galapagos, Vulnerable species
Dalmation dogs playing on beach, Norfolk, UK, August
Atlantic grey seal (Halichoerus grypus) resting vertical beneath the surface, Lundy Island, Devon, UK. July
Pink abalone (Haliotis corrugata) shell detail, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, January
Lush vegetation and traditional statues in the Marquesas Islands, French Polynesia
Adelie penguins (Pygoscelis adeliae) on iceberg, Antarctica, January
Rain shower approaching La Vieille lighthouse, Pointe du Raz, Brittany, France. November 2006
Flock of Sanderling (Calidris alba) flying over a rough sea, Berwickshire Coast, Scotland, UK, March
The Long Wall, aerial view of sand dunes bordering the atlantic coastThe " Long Wall", aerial view of sand dunes bordering the atlantic coast, near Swakopmund, Namib desert, Namibia, August 2008
Aerial view of rivers and mangrove forest. Sarawak Mangrove Reserve, Sarawak, Borneo, Malaysia. June 2006
Hartland Quay, rugged north Devon coastline, nr Hartland, Devon, UK. March 2009
Aerial view of Jersey, Channel Islands. May 2009
Grey seal {Halichoerus grypus} two adolescents interact among the breaking waves, Lincolnshire, UK
Aerial view of the Rock Islands, Palau, Micronesia, December 2001
King Penguins (Aptenodytes patagonicus) head portraits, on the beach at Salisbury Plain, South Georgia, South Atlantic. January
Emperor Penguins walking in a row, Antarctica
Sea and stgones at Porth Nanven, Cot Valley, near Cape Cornwall, West Cornwall, UK. June 2012
Hartland Quay, in late evening light and rising tide, Devon, UK. May 2012
Whitsunday Islands, aerial view, Great Barrier Coral Reef, Queensland, Australia, October 2011
Fulmar (Fulmarus glacialis) rear view of bird hanging in air over steep cliffs, Shetland Islands, Scotland, UK. HIGHLY COMMENDED - Animals in their environment - Wildlife Photographer of the Year
Baby Olive Ridley sea turtle (Lepidochelys olivacea) struggles against the swell to swim away from the beach where it hatched, Ostional, Costa Rica. October
View across Oxwich Bay on the Gower Peninsula, South Glamorgan, Wales, January 2012
Sailing yacht at sunset off coast of Hanalai Bay, Kauai, Hawaii, USA
Herreshoff S-Boat "Sturdy Beggar" built in 1922 and restored by the International Yacht Restoration School (IYRS) in Newport, Rhode Island, USA
Little ringed plover {Charadrius dubius} nest with four eggs camouflaged on shingle, Lorraine, France
Grey seal (Halichoerus grypus) on stone beach, yawning, Saltee Islands, County Wexford, Ireland, May
Polar bear (Ursus maritimus) spring cub resting on newly formed pack ice, off the 1002 area of the Arctic National Wildlife Refuge, Alaska, USA, October
Aerial view to the west of Portbraddan on the north coast of County Antrim, Northern Ireland, UK, September 2009
Flints on West Runton beach, Norkfolk, UK July
Sunset views of La Corbiere lighthouse located at the extreme south-western point of Jersey, St. Brelade) Jersey, The Channel Islands, UK, September 2011
Moon over Cathedral Cove at dawn, near Hahei, Coromandel Peninsula, North Island, New Zealand. December 2006
