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Alcedo giant tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island. This is where the largest extant population exists
Brown pelican (Pelecanus occidentalis) landing reflected in tide pool, Puerto Egas, Santiago Island, Galapagos
Galapagos hawk (Buteo galapagoensis) in flight, Alcedo Volcano, Isabela Island, Galapagos
Wildlife on the coast of Cape Hammond including Galapagos flightless cormorant (Phalacrocorax harrisi), Sally lightfoot crab (Grapsus grapsus) and Marine iguana (Amblyrhynchus cristatus)
Landscape of Bartolome Island at sunset, Galapagos
Galapagos Sea Lion (Zalophus californianus wollebacki) resting on rocks at sunset on North Seymour Island, Galapagos, Ecuador. December
Luilaka River with clouds reflected in the surface of the water, Salonga National Park is on the left bank. Democratic Republic of the Congo
Granite boulders in Spitzkoppe mountains, Namib Desert, Namibia, October
Coastal landscape with empty beach Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, July
Split level image of a Southern stingray (Dasyatis americana) swimming over a sand bar. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Light patterns on the seabed and fluffy cumulus clouds in the sky in a split level image. North Sound, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Mountains, Nubra Valley, Karakorum Wildlife Sanctuary, Ladakh, India. September 2011
Coast of l Ile-des-Pins with the New Caledonia pines (Araucaria columnaris) that gave the name to the island at sunset, New Caledonia, Pacific Ocean
Stormy sky over Klein Schietveld, Brasschaat, Belgium. March 2019
Guanacos (Lama guanicoe), five silhouetted on slope in evening. Torres del Paine National Park, Patagonia, Chile. December
Peaks and clouds above Tiger Leaping Gorge at dawn. Three Parallel Rivers of Yunnan Protected Areas World Heritage Site, Yunnan Province, China. 2010
Cley Mill and Reedbed in winter snow storm, Norfolk, UK, February
Common Hawthorn (Crataegus monogyna) tree growing out of joints in Carboniferous limestone, Ingleton, Yorkshire, January 2017
Pine trees in mist, Ballons des Vosges Regional Natural Park, Vosges Mountains, France, October 2014
Autumnal woodland and Young Peak surrounded by clouds, British Columbia, Canada, September 2013
A misty morning over a mixed woodland in autumn. Kinnoull Hill Woodland Park, Perthshire, Scotland, October 2012
Llyn (Lake) Padarn looking looking souh-east towards Llanberis with two Mute Swans (Cygnus olor) swimming Snowdonia National Park, North Wales, UK, September 2017
Pine forest and mountains reflected in Loch Morlich, Rothiemurchus, Cairngorms National Park, Scotland, UK, September 2016
Loch Lurgainn and Stac Pollaidh from Cul Beag, at sunset, Assynt, Highlands, Scotland, UK, September 2013
View from summit of Cul Beag overlooking Loch Lurgainn and Stac Pollaidh at sunset, Coigach, Scotland, September
Route One, Southeast Iceland. December
View of Rutland Water Nature Reserve, RAMSAR site, Special Protection Area and Site of Special Scientific Interest, August 2016, Egleton, Rutland, England
Reindeer (Rangifer tarandus) silhouetted against sky, walking across upland, Cairngorms, Scotland, UK, August
RF- Yellow Labrador retriever sitting in cornfield, UK, August
Moriche plam tree (Mauritia flexuosa) at Laguna Grande in an area of flooded forest, Cuyabeno wildlife reserve, Sucumbios, Amazon rainforest, Ecuador, July, Cuyabeno wildlife reserve, Sucumbios
The Melnik Earth Pyramids, rock formations, known as hoodoos, situated at the foothills of the Pirin mountain range in south-western Bulgaria, Bulgaria, April
The Quiraing bathed in morning light, eastern face of Meall na Suiramach, the northernmost summit of the Trotternish on the Isle of Skye, Scotland, UK. November 2017
RF- New Forest pony near Bratley View, New Forest, Hampshire, UK. November
Bottle (Battle) Mountain, a granite outcrop in South Rupununi savanna, Guyana, South America
Mountain hare (Lepus timidus) three adults on a snow slope high in the Cairngorm Mountains, Cairngorms National Park, Scotland, UK, February
Zebra at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017
English oak tree (Quercus robur) in field of Oilseed rape / rapeseed (Brassica napus), UK, May 2017
Blue wildebeest (Connochaetes taurinus), migrating herd silhouetted on the horizon at sunset, Masai Mara Game Reserve, Kenya
Mount Lindesay, forests and low clouds at sunset, view from Kamarun lookout. Green Mountains, Lamington National Park, Rainforests of Australia UNESCO World Heritage Site, Queensland, Australia
Fort Collins- Colorado forest fire seen from Estes Park. The fire was intensified by dead trees killed by the Mountain pine beetle (Dendroctonus ponderosae) Colorado, USA, June
African elephants (Loxodonta africana) grazing on shore of Lake Kariba, Matusadona National Park, Zimbabwe
Mountain forests where the Yunnan snub-nosed monkey (Rhinopithecus bieti) lives, Yunnan, China
Blacktip reef shark (Carcharhinus melanopterus) just at the surface off the island of Yap, Micronesia
Cley Beach with scattering of light snow, and boats, Norfolk, England, UK, December 2010
Cloud forest scene in the Monteverde Cloud Forest Reserve, Costa Rica, February 2015
Bird nesting cliffs at Haeavikurbjarg, Hornvik, Hornstrandir, Iceland. July
Thonga Beach on the coast of Indian Ocean, Maputuland, KwaZulu-Natal, South Africa, January
Mountainous landscape in Torres del Paine National Park, with mountains behind, Chile, June 2015