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Split level image of a Southern stingray (Dasyatis americana) swimming over a sand bar. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Oceanic whitetip shark (Carcharhinus longimanus) is framed by the surface. Elphinstone reef, Marsa Alam, Egypt. Red Sea
Spotted eagle rays (Aetobatus narinari) through a large window on the wreck of the former mine sweeper C-56 Juan Escutia. Cancun, Mexico
A split level photo of a female southern stingray (Dasyatis americana) swimming over seabed. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Dusky Shark (Carcharhinus obscurus) solitary shark patrols the deep blue waters off Port St Johns, South Africa
Oceanic whitetip shark (Carcharhinus longimanus) swimming close to surface. Brothers Islands, Egypt
Caribbean reef shark (Carcharhinus perezi) patrolling coral reef with Schoolmaster snapper (Lutjanus apodus) and other fish. Caribbean Sea off Gardens of the Queen National Park, Cuba
Silky shark (Carcharhinus falciformis), three in Caribbean Sea off Gardens of the Queen National Park, Cuba
Angel shark (Squatina squatina) laying on the sand, Canary Islands
Reef manta (Mobula alfredi) female swimming close to a coral reef, while cleaner wrasse (Labroides dimidiatus), tiny by comparison, pick parasites from her lips
Blue shark (Prionace glauca) off Halifax, Nova Scotia, Canada. July
A close up of a blue shark (Prionace glauca) as it investigates the camera beneath the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August
Blue shark (Prionace glauca) Azores, Atlantic Ocean
Nurse sharks (Ginglymostoma cirratum) three in a courtship dance at sunrise in a mangrove area near Eleuthera, Bahamas
Whale shark (Rhincodon typus) front view portrait, Cenderawasih Bay, West Papua. Indonesia
Whale shark (Rhincodon typus) near surface, viewed from above, Cenderawasih Bay, West Papua. Indonesia
A Caribbean reef shark (Carcharhinus perezi) just below the surface. Split level with blue sky and clouds. Long Island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean
Silky shark (Carcharhinus falciformis) San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve, Socorro Islands, Western Mexico
Great hammerhead shark (Sphyrna mokarran) swimming over sandy seabed, South Bimini, Bahamas. The Bahamas National Shark Sanctuary, West Atlantic Ocean
Nurse shark (Ginglymostoma cirratum) throwing up sand as it hunts in the sandy seabed, South Bimini, Bahamas. The Bahamas National Shark Sanctuary, West Atlantic Ocean
Blacktip reef shark (Carcharhinus melanopterus) just at the surface off the island of Yap, Micronesia
Mobula / Chilean devil ray (Mobula tarapacana) with a school of fish (Capros aper) showing vertical migration, Santa Maria, Azores, Portugal, Atlantic Ocean
River stingray (Potamotrygon sp) Formoso River, Bonito, Mato Grosso do Sul, Brazil
Split level view of Blue shark (Prionace glauca) at surface at sunset, Azores Islands, Portugal, Atlantic Ocean
Spotted eagle ray (Aetobatus narinari) swimming in the blue next to a coral reef. Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Blacktip reef shark (Carcharhinus melanopterus) swimming just below surface at sunset, split level view. Off the island of Yap, Micronesia
Great white shark (Carcharodon carcharias) dorsal fin visible on surface. Guadalupe Island, Mexico
Undulate Ray (Raja undulata) Sark, British Channel Islands, July
Blonde Ray (Raja brachyura) demonstrating camouflage against sandy habitat. Channel Islands, UK, August
Basking shark (Cetorhinus maximus) feeding, Mull, Scotland, June 2009
Two Basking sharks (Cetorhinus maximus) Mull, Scotland, June 2009
Basking shark (Cetorhinus maximus) Mull, Scotland, June 2009
Round stingray (Taeniura grabata) swimming over rocks, Princesa Alice, Azores, Portugal, June 2009
Silvertip sharks (Carcharhinus albimarginatus) in open water, Revillagigedo islands, Mexico. Pacific Ocean
Whaleshark (Rhincodon typus) swimming and filtering fish eggs from the water. A behaviour known as ram-feeding. Isla Mujeres, Quintana Roo, Yucatan Peninsular, Mexico. Caribbean Sea
Blue shark (Prionace glauca) female off the coast of Portugal. August
Large female Southern stingray (Hypanus americanus) in shallow water, under storm clouds, Stingray City, Grand Cayman. Stingray City is feeding site for stingrays and bait was used on this shoot
Portrait of Whale shark (Rhincodon typus) at the surface. Isla Mujeres, Quintana Roo, Yucatan Peninsular, Mexico. Caribbean Sea
Blue shark (Prionace glauca) near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August
Blue shark (Prionace glauca) swimming near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August
Caribbean reef sharks (Carcharhinus perezi) swimming through the middle of a school of yellowtail snappers (Ocyurus chrysurus). Grand Bahama, Bahamas. Tropical West Atlantic Ocean
RF - Silky shark (Carcharhinus falciformis) swimming beneath the surface of the ocean, with evening sun burst. Gardens of the Queen National Park, Cuba
Caribbean reef shark (Carcharhinus perezi) swimming over Common sea fans (Gorgonia ventalina) on a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Whale shark (Rhincodon typus) mouth open feeding at the surface, Isla Mujeres, Yucatan Peninsular, Mexico. Caribbean Sea
Great hammerhead shark (Sphyrna mokarran) portrait, South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
RF - Nurse sharks (Ginglymostoma cirratum) pair swimming over sand in shallow water. White spiracle behind eye is visible. South Bimini, Bahamas. The Bahamas National Shark Sanctuary
A long exposure of a Tiger Shark (Galeocerdo cuvier) portrait, at the surface at dusk. Little Bahama Bank, Bahamas. West Atlantic Ocean. March