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Female Grizzly bear (Ursus arctos horribilis) standing up, with two cubs nearby, Khutzeymateen Grizzly Bear Sanctuary, British Columbia, Canada, June
Tree sparrows (Passer montanus) perching on a branch in the rain. Perthshire, Scotland, December
Tabby cat, Max, 5 months, with smoke black kitten
Chacma baboon (Papio hamadryas ursinus) portrait of adult male. deHoop Nature reserve. Western Cape, South Africa
Grey heron (Ardea cinerea) adult in breeding plumage, close-up of head and colourful orange beak, backlit, Berlin Zoological Garden, Berlin Zoo, Germany. February
Lion (Panthera leo) male shaking his head after rain, Masai-Mara Game Reserve, Kenya
Starfish Beach, with many starfish in the shallow sea (Asteroidea) Colon Island, Bocas del Toro Archipelago, Bocas del Toro Province, Panama
White-tailed eagle (Haliaeetus albicilla) in flight, Norway, April
Alpaca Herd on Norfolk Farm, winter
Polar bear (Ursus maritimus) portrait, female. Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Lesser flamingo (Phoeniconaias minor) portrait, Lake Bogoria, Rift valley, Kenya, Africa
Topi (Damaliscus korrigum) at sunset, Masai-Mara Game Reserve, Kenya
Portrait of brown and white coated Holland lop-eared domestic rabbit (Oryctolagus cuniculus domesticus), Lorraine, France, July
Large group of Cavalier King Charles Spaniel, puppies, three of which have tricolour coat and the rest blenheim
French Bulldog bitch with blue-white, aged one year
Nosy Be Pygmy Leaf Chameleon (Brookesia minima) female, worlds smallest reptile. Nosy Be, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, January 2010
Indri (Indri indri) calling, Andasibe-Mantadia NP, Madagascar
Fresh Wych elm (Ulmus glabra) leaves in spring. Dorset, UK May
Lion(Panthera leo) cub playing with its mothers tail, Masai-Mara Game Reserve, Kenya. Vulnerable species
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Gemsbok (Oryx gazella) silhouetted at dawn, Kalahari Desert, Botswana
Lesser flamingo flock (Phoeniconaias) flying over lake with green algae, aerial view, Rift valley, Kenya
Bramble (Rubus plicatus) leaves and stem against white background. France, August
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Black-headed gull (Chroicocephalus ridibundus) alighting on water at dawn, Cheshire, UK, November. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Grants zebra (Equus burchelli boehmi) herd with large cloud above, Amboseli National Park, Kenya
Aerial view of the River Kinabatangan and riverine tropical rainforest, Sabah, Malaysia, Borneo, Asia
Giant deep-sea isopod (Bathynomus giganteus) Picture was taken in cooperation with the Zoological Museum University of Hamburg
Toco Toucan (Ramphastos toco) beak open with tongue visible while feeding on mango, Brazil
Atlantic Puffin (Fratercula arctica) in flight, coming into land with beak full of sand eels (Ammodytes tobianus). Fair Isle, Shetland Islands, Scotland, UK, July
Black howler monkeys (Alouatta caraya) male and two females calling from tree, captive, Apenheul Park, Netherlands
Polar bear (Ursus maritimus) juvenile rolling around on newly formed pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Palm (Areaceae) leaves silhouetted in mist at dawn, Royal Chitwan National Park, Nepal
People watching the Fissure eruption of Chico volcano into 9km diameter caldera. Sierra Negra volcano, Isabela Island, Galapagos, Ecuador. October 2005
Okarito Brown Kiwi (Apteryx rowi) feather detail resembling mammalian hair. Okarito Forest, Westland, South Island, New Zealand
Detail of small waves on the eastern Atlantic. Isle of Coll, Scotland, October 2012
Great white egrets (Casmerodius albus) reflected in Pulicat Lake, Tamil Nadu, India, January 2013
Fireflies {Lampyridae} in rainforest, Singapore
Aerial view of Lago Preto, Amazon Rainforest and lake, Peru
A Kamchatka Brown Bear (Ursus arctos beringianus) standing in breaking waves. Pacific coast of Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Japanese Macaque (Macaca fuscata) juvenile appears to be doing tricks to get some attention in Jigokudani, Japan
An Einsiedler / Swiss warmblood mare (Equus caballus) cantering, Schwyz, Switzerland, July
Roe Deer (Capreolus capreolus) in woodland, Peerdsbos, Brasschaat, Belgium, June
Black-headed Ibis / Oriental Ibis (Threskiornis melanocephalus), Karnataka, India
Asian Elephant (Elephas maximus) femalenext to termite mound in forest, Nagarhole National Park, South India
Devils Marbles, Devils Marbles Conservation Reserve, Northern Territory, Australia, June 2010
Spotted hyena (Crocuta crocuta) hunting Lesser flamingoes (Phoenicopterus minor) Lake Nakuru, Kenya
Black kitten in garden, Germany
