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Adult male Hippopotamus (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November
Australian Sea Lion (Neophoca cinerea) sitting on beach with one flipper up, Seal Bay Conservation Park, Kangaroo Island, South Australia State, Australia
Polar bear (Ursus maritimus) curious cub sits up on its hind legs, paw raised, and tries to balance itself, along Bernard Spit in autumn, 1002 area of the Arctic National Wildlife Refuge, Alaska
Zebra (Equus quagga) foal running through Devils Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January
Detail of plumage of Great White Egret / Heron (Ardea herodia occidentalis). Ding Darling Reserve, Florida, January
Sleeping mixed-breed piglet in straw, Maple Park, Illinois, USA
Tranarossan Bay, near Melmore Head, Rosguill Peninsula, County Donegal, Republic of Ireland, August 2011
Polar bear (Ursus maritimus) curious adult investigates a remote camera along a hillside on Spitsbergen and the northwest coast of the Svalbard Archipelago, Norway, Greenland Sea, July
Arctic Fox (Vulpes lagopus semenovi), in dark summer pelage, patrolling cliffside rocks along the coast of Medny Island in the Commander Islands (aka Komandorsky Islands), Russia, July
Remarkable Rocks at night, Flinders Chase National Park, Kangaroo Island, South Australia State, Australia, September 2011
Indian moon / Indian luna Moth (Actias selene) close up of defensive spines on caterpillar. Captive
Mountain Hare (Lepus timidus) on moorland with white winter coat, Kinder Scout, Peak District National Park, Derbyshire, UK, February
Common starling (Sturnus vulgaris) flock at dusk engaging in pre-roost display, Dumfries-shire, Scotland, March
Brain coral (Diploria strigosa) photographed under blue light to excite the fluorescence, Little Cayman, Cayman Islands, British West Indies, Caribbean Sea
Scots pine (Pinus sylvestris) mature tree in evening light, Abernethy Forest RSPB Reserve, Cairngorms National Park, August
Sandstone cliffs, Rackwick Bay, Hoy, Orkney, Scotland, July 2004
Mustang / wild horses, second year cremello colt Claro play fighting with bay, McCullough Peak herd, Wyoming, USA, July 2009
Grey angelfish (Pomacanthus arcuatus) portrait, Georgetown, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Rock beauty angelfish (Holacanthus tricolor) pair at dusk performing their courtship dance with the larger male circling the female, Georgetown, Grand Cayman, Cayman Islands, British West Indies
Schoolmaster snapper (Lutjanus apodus) above a coral reef, Georgetown, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Western grebe (Aechmophorus occidentalis) pair swimming, with reflections, Bolsa Chica Ecological Reserve, California, USA, February
Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February
Kangaroo Ridge in the North Cascades, in winter, Okanogan National Forest, Washington, USA, November 2010
Chilean huemul or South Andean deer (Hippocamelus bisulcus), Torres del Paine National Park, Patagonia, Chile. Endangered species
Briard / Berger de Brie, head portrait with plaited hair between eyes
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched together)
African lions (Panthera leo) one lioness licking another, Okavango Delta, Botswana
Gloucester old spot piglets (Sus scrofa domestica) suckling at the Cotswold Farm Show, Cirencester, UK, July 2008
Rothschilds giraffe (Giraffa camelopardalis rothschildi) close up of young calf skin pattern, captive
Grevy's zebra (Equus grevyi) close up, captive
Red fox (Vulpes vulpes) close-up of half of face, captive
Black bear (ursus americanus) cub standing on hind legs, Yellowstone National Park, USA, June
Grizzly Bear (Ursus arctos horribilis) male in water. Coastal Katmai National Park, southwest Alaska, USA, August
Blue Shark (Prionace glauca) near sea surface and divers in background. Santa Maria, Azores, September
Chinese Crested Dog, hairless bitch with puppy, male, 4 months
Male African Lion (Panthera leo) playing with 4 month cub. Big Marsh, near Ndutu, Nogorongoro Conservation Area, Serengeti National Park, Tanzania
Demoiselle crane (Anthropoides virgo) portrait, captive
Icebergs off the Antarctic Peninsula, Antarctica, February 2009, Taken on location for BBC Frozen Planet series
Corfe Castle rising out of mist, viewed from Kingstone, Purbeck, Dorset, UK, September 2010
Beech trees in autumn mist, Morvan regional park, Burgundy, France, November 2010
Aphelandra (Acanthus sp) flowering. Oahu, Hawaii, February
Common buckeye butterfly (Junonia coenia) feeding on Ironweed (Vernonia altissima) flower, North Guilford, Connecticut, USA, August
Gulls on the beach at sunset, The Wash, Hunstanton, Norfolk, UK, September 2011
Leatherback Turtle (Dermochelys coriacea) returning to the sea after egg laying. Cayenne, French Guiana, June
Male Greenfinches (Carduelis chloris) squabbling in flight. Dorset, UK, March
Ocellated eagle ray (Aetobatus ocellatus), underside, Natal Coast, South Africa
Black blotched pufferfish (Diodon liturosus) inflated with water at night as defensive behaviour, Sipadan Island, Borneo, Malaysia
Lone Maple (Acer) in a field against cumulo-nimbus and mammatus clouds. Picardy, France, May
