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Brown bear (Ursus arctos) and Grey wolf (Canis lupus) together in wetlands, Kuhmo, Finland, July
King penguins (Aptenodytes patagonicus) crossing water to reach breeding site, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Flemish giant rabbit with ears erect
Leatherback Turtle (Dermochelys coriacea) hatchling crossing a beach to get to the sea. Cayenne, French Guiana, July
Northern / Hen Harrier (Circus cyaneus) and Kestrel (Falco tinnunculus) below, fighting in flight. Vosges, France, February
Volcanic eruption, Eyjafjallajokull, near the Myrdalsjokull glacier, South Iceland, april 2010
Portrait of a young Little Owl (Athene noctua)
Portrait of English Springer Spaniel (field type). Elkhorn, Wisconsin, USA, January
Tricolour Border Collie puppy
Row of Larch trees on flowering heather moorland, Lammermuir Hills, Berwickshire, Scotland, August
Killer Whales (Orcinus orca) spy-hopping to observe Weddell Seal (Leptonychotes weddellii) in preparation to knock it from the ice by creating a wave. Marguerite Bay, Antarctic Peninsula, summer
Female Scottish wildcat (Felis silvestris grampia) portrait, in winter coat, semi habituated living wild in native oakwood, Lochaber, Scotland, April
Atlantic puffin (Fratercula arctica) gathering around burrows on cliff top, Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland, UK, June
Kilchurn Castle on Loch Awe, Argyll and Bute, Strathclyde, Scotland, UK, April 2011
Emergent Menggaris Tree / Tualang (Koompassia excelsa) protruding the canopy of lowland rainforest. Danum Valley, Sabah, Borneo
Adult Adder (Vipera berus) basking on pebble beach. Abbotsbury, Dorset, UK, March
Storm clouds building up off coasts of Norfolk, Weybourne Cliffs, Norfolk, UK, November
Detail of frost on Western Red Cedar (Thuja plicata). Vancouver Island, Canada, March
Colourful water-smoothed pebbles on shingle beach. Normandy, France, October 2010
Wild Horses / mustangs, two stallions rearing up fighting, Pryor Mountains, Montana, USA, June
Pink abalone (Haliotis corrugata) shell detail, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, January
Emperor penguins (Aptenodytes forsteri) explode out of the water, returning to breed at Cape Washington, Antarctica. Taken on location for BBC Frozen Planet series, 2009
Boomslang (Dispholidus typus) Immature male hunting in thorn bush above deHoop Vlei, Western Cape, South Africa, December
King Eider (Somateria spectabilis) flying low over water. Floe edge, Arctic Bay, Nunavut, Canada, April
Moon rising over winter landscape, stubble field and Oak trees, Gimingham, Norfolk, UK
Mixed flock of Snow geese (Chen caerulescens atlanticus / Chen caerulescens) and Sandhill cranes (Grus canadensis) at wintering area at dawn, with full moon, Bosque del Apache, New Mexico, USA
Border Collie puppy, 13 weeks
Golden Retriever puppy sitting in snow in late afternoon. Big Rock, Illinois, USA, February
Two smooth coated Jack Russell Terrier puppies, tan and white, 6 weeks
Rough coated Jack Russell Terrier puppy, black and tan, portrait
Common Puffball fungus (Lycoperdon perlatum) emitting spores into the air
Jersey tiger moth (Euplagia quadripunctaria) large numbers resting on trunk of Sweetgum tree (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011
Abstract image of large sea swells hitting the rocks, Portuguese southwestern coast, Portugal, March, 2010
Gorgonian Seafan Coral (Gorgonacea) polyps open to feed at night. South China Sea, Sipadan Island, Borneo
Bullseye / Io moth (Automeris io) showing eye spot markings on wings during deimatic display to deter predators, photographed on a white background, originating from North and Central America
Indian moon / Indian luna moth (Actias selene) head-on view showing feather-like antennae. Captive
Path towards the coastguard cottages and the Seven Sisters Chalk Cliffs. Seaford Head Nature Reserve, East Sussex, UK, May 2009
Portrait of a Birman-cross cat
Silver tabby Chinchilla male cat
Black-and-white Border Collie bitch lying chin on floor with black-and-white kitten
Cairns birdwing butterfly (Ornithoptera priamus) male on fern leaf and Blue Mountain Swallowtail butterfly in flight, Atherton Tablelands, Queensland, Australia, October
Noctule bat (Nyctalus noctula) Captive, UK
Flock of Common Starling (Sturnus vulgaris) gathering before landing at winter roost, Salisbury Plain, Wiltshire, UK, January
Horse in snow storm with shed in background, USA
Wild Horse / Mustang shaking head and mane, Adobe Town Herd Area, southwestern Wyoming, USA, WINNER OF WILD HORSE SECTION, EQUINE IDEAL PHOTOGRAPHY CONTEST 2012
Mountain pansy (Viola lutea) flowers, Strathspey, Cairngorms National Park, Scotland, June
Cliff scenery at North Hoy, Hoy, Orkney, Scotland, July 2008
Feral Goat (Capra hircus) male, Inverness-shire, Scotland, July