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Head portrait of Bornean Horned Frog (Megophrys nasuta) among the leaf litter in the lowland rainforest of Borneo. Danum Valley Conservation Area, Sabah, Malaysia
Arctic fox (Alopex lagopus) in winter coat, walking across snow, Svalbard, Norway. April
Ptarmigan (Lagopus mutus) in winter plumage, resting in snow, Svalbard, Norway. April
Polar bear (Ursus maritimus) female standing on snow with cub peering out behind, Svalbard, Norway. April
Polar bear (Ursus maritimus) female with cub standing on ice, Svalbard, Norway. April
Polar bear (Ursus maritimus) female, resting on ice with cub sitting next to her waving and poking out tongue, Svalbard, Norway. April
Sand cat (Felis margarita) walking over sandy ground. Captive, occurs in North Africa, the Arabian Peninsula, Pakistan and the Middle East
Agile frog (Rana dalmatina) sitting in autumn leaves, Upper Bavaria, Germany. October
Santa Catalina Island rattlesnake (Crotalus catalinensis), only rattlesnake without rattle, slithering in rock. Santa Catalina Island, Loreto Bay National Park, Sea of Cortez, Mexico. May
Agile frog (Rana dalmatina) sitting in autumn leaves on forest floor, Upper Bavaria, Germany. October
Cheetah (Acinonyx jubatus) female, walking through long grass at sunrise, Ndutu area, Serengeti / Ngorongoro Conservation Area, Tanzania
Wolf (Canis lupus) walking in forest, Finland. September
Common ghostgoby (Pleurosicya mossambica) camouflaged on a hard coral colony (Plesiastrea sp)Common ghostgoby (Pleurosicya mossambica) camouflaged on a hard coral colony (Plesiastrea sp.), Ambon Bay, Maluku Archipelago, Indonesia. Banda Sea
Tawny owl (Strix aluco) roosting in Silver birch (Betula pendula) tree in daytime, Cairngorms National Park, Scotland, UK
Deep red Feather star shrimp (Hippolyte prideauxiana) crawling secretly amongst the arms of a Feather star (Antedon bifida), Loch Duich, Highands, Scotland, UK
Two Pallas's cat (Otocolobus manul) kittens waiting at their den for their mother to return from hunting, Mongolia, June, 2017. Filmed for BBC series Big Cats
A female Pallas's cat (Otocolobus manul) with two young kittens hiding amongst rocks at their den site, Mongolia, June, 2017. Filmed for BBC series Big Cats
Minor's chameleon (Furcifer minor) on tree branch, Madagascar
Leafy seadragon (Phycodurus eques) demonstrates the effectiveness of its camouflage as it shelters amongst seaweeds. This individual is a male carrying eggs
European rabbit (Oryctolagus cuniculus) in steppe habitat. Alfaro, La Rioja, Spain. Endangered species
Pair of Coleman shrimps (Periclimenes colemani) living in a fire urchin (Asthenosoma varium). The female is the larger shrimp
Great grey owl (Strix nebulosa) camouflaged in birch woodland, Finland, March
Willow grouse (Lagopus lagopus) camouflaged against snow, Inari, Kiilopaa, Finland, February
Lioness (Panthera leo) and cubs in rain, Masai Mara Game Reserve, Kenya, September
Knysna turaco (Tauraco corythaix) in tree, Kariega Game Reserve, Eastern Cape, South Africa
Ornate ghost pipefish (Solenostomus paradoxus) young hides amongst the branches of a black coral bush (Antipathes sp) Sangeang Island, Sumbawa, Indonesia, Flores Sea
Little ringed plover {Charadrius dubius} nest with four eggs camouflaged on shingle, Lorraine, France
Grey seal (Halichoerus grypus) on stone beach, yawning, Saltee Islands, County Wexford, Ireland, May
Cheetah at dawn {Acinonyx jubatus} Masai Mara, Kenya
Arctic fox male portrait {Vulpes lagopus} Norway captive
Warty frogfish (Antennarius maculatus) yawning in its hiding place amongst white sponges, Bitung, North Sulawesi, Indonesia, Molucca Sea
Leopard (Panthera pardus) hiding in dense foliage, Bardia National Park, Terai, Nepal
Snowy owl (Bubo scandiaca) standing behind dried plant in snow, Canada. January
Orange tip butterfly (Anthocharis cardamines) caterpillar crawling over Garlic mustard (Alliaria petiolata) leaf, Orvieto, Umbria, Italy. April
Snowy owl (Bubo scandiaca) female, in flight in winter, Canada. January
Female Polar bear (Ursus maritimus) with cub in snow, Churchill, Canada. November
Arctic hare (Lepus arcticus groenlandicus) portrait, Ittoqqortoomiit, Northeast Greenland. February
Lioness (Panthera leo) lying in long grass on the savannah, portrait, Okavango Delta, Botswana, Africa
Painted frogfish (Antennarius pictus) portrait, Sipidan, Malaysia, Celebes Sea
Long-snouted seahorse / Spiny seahorse (Hippocampus guttulatus) camouflaged on a White seafan (Eunicella singularis), Marine Protected area Punta Campanella, Costa Amalfitana, Italy, Tyrrhenian Sea
Male Jackson's chameleon (Chamaeleo jacksoni) reaching out as it moves between leaves, Maui, Hawaii
Dense growth of Bootlace seaweed (Chorda filum) in shallow water, with a Stalked jellyfish (Calvadosia sp). clinging on between strands, Kimmeridge, Dorset, England, UK, English Channel. July
Female polar bear (Ursus maritimus) with two cubs walking in a line across snow, Churchill, Canada. November
Polar bear (Ursus maritimus) sitting in snow during a blizzard, Churchill, Canada. November
An endemic Hawaiian pom-pom crab / Boxer crab (Lybia edmondsoni) carrying a juvenile Anemone (Triactis producta) in its claws as a defense mechanism
Two Mountain hares (Lepus timidus) running in deep snow, Monadhliath Mountains, Highlands, Scotland, UK. March
Mountain hare (Lepus timidus) running over snow on hillside, Monadhliath Mountains, HIghlands, Scotland, UK. February
Mountain hare (Lepus timidus) running down a snowy mountainside, Monadhliath Mountains, HIghlands, Scotland, UK. February