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Red-and-green macaw (Ara chloropterus) in flight over forest canopy. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
Two Jaguars (Panthera onca) snarling at each other on the riverbank, Pantanal, Brazil. September
Toco Toucan (Ramphastos toco) head and beak profile portrait, Brazil
Blue iridescent scales of a Nymphalid butterfly (Epiphile orea) magnified 11x, deceased insect found in Piedade, Sao Paulo, Brazil. South-east Atlantic Forest
Hyacinth macaws (Anodorhynchus hyacinthinus) mutual preening, Estrada Parque, Pantanal, Brazil
Burrowing owl (Athene cunicularia) fledgling with head on side, Pantanal, Brazil. November
Green-winged macaws (Ara chloroptera) preening each other. Brazil. South America
Toco Toucan (Ramphastos toco) beak open with tongue visible while feeding on mango, Brazil
Sequence of a Red-and-green macaw or green-winged macaw (Ara chloropterus) coming into land at its nest hole on a cliff. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
RF - Red-and-Green Macaws or Green-winged Macaws (Ara chloropterus) in flight over forest canopy. Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Pink Ipe trees (Tabebuia ipe / Handroanthus impetiginosus) in flower, Pantanal, Mato Grosso State, Western Brazil
Maned wolf (Chrysocyon brachyurus) setting out to hunt in grassland at dusk. Serra da Canastra National Park, Brazil
Capybara (Hydrochoerus hydrochaeris) swimming, Little Paraguay River, Pocone, Brazil
Spear-nosed bat (Phyllostomus elongatus) hanging from a tree branch, Manaus, Brazil
Common vampire bat (Desmodus rotundus), portrait, Pantanal, Mato Grosso Brazil
RF - Red-and-green macaws (Ara chloropterus) two with one taking off, Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Red-and-green macaw (Ara chloropterus) in flight, Pantanal, Brazil. August
Golden-headed lion tamarin (Leontopithecus chrysomelas) feeding on jackfruit, coastal rainforest, Mata Atlantica, Bahia, Brazil
Giant Otters (Pteronura brasiliensis) running along a sand bar. Cuiaba River, Pantanal, Brazil
White-cheeked spider monkey (Ateles marginatus), Amazon, Brazil. June
Hyacinth Macaw (Anodorhynchus hyacinthinus) adult in nest hole in tree, hole in a tree, Pantanal area, Mato Grosso, Brazil
Floating leaves of Giant water lily (Victoria amazonica). Northern Pantanal, Cuiaba River, Mato Grosso, Brazil
Preening red-and-green macaw or green-winged macaw (Ara chloropterus) (Family Psittacidae). Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Yellow Anaconda (Eunectes notaeus) in vegetation at the edge of the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Yacare caiman (Caiman yacare) Cuiaba River, Pantanal, Brazil, August
Hyacinth Macaw (Anadorhynchus hyacinthinus) two playing whilst hanging upside down, Pantanal, Brazil
Two Giant Otter / Giant Brazilian Otter (Pteronura brasiliensis) sunbathing on a tree trunk. The Pantanal wetlands of Mato Grosso State, Brazil, October
Jaguars (Panthera onca) courting pair on a sand bank. Cuiaba River, Northern Pantanal, Mato Grosso, Brazil
Piraputanga, (Brycon hilarii) reflected on the water surface, Aquario Natural, Bonito, Mato Grosso do Sul, Brazil
Jaguar (Panthera onca) Pantanal, Mato Grosso, Brazil
Grass fire at night in Pantanal, Brazil
Julia heleconia (Dryas iulia) butterfly on head of Yacare caiman (Caiman yacare). Butterflies often land on caimans head to drink the salt from its eyes. Pantanal, Brazil
Spectacled caiman (Caiman crocodilus) in water at night, Mato Grosso, Pantanal, Brazil
Jaguar (Panthera onca palustris), male hunting amongst tree roots at edge of the Cuiaba River. Three Brothers, Porto Jofre, northern Pantanal, Mato Grosso, Brazil. August 2017
Marsh deer (Blastocerus dichotomus), stag. Pousada Araras, northern Pantanal, Mato Grosso, Brazil
Sequence of a Three-banded armadillo (Tolypeutes tricinctus) unrolling from defensive ball posture to walking. Captive
Yacare caiman (Caiman yacare) group basking, mouths open to keep cool. Pantanal Matogrossense National Park, Pantanal, Brazil
Yacare caiman (Caiman yacare) feeding on Amazon sailfin catfish (Pterygoplichthys pardalis) Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Yacare caiman, (Caiman yacare) Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) male on riverbank, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) female, smelling scent marking of male. Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) on river bank, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) male drinking, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) male, Pantanal, Brazil
Female Jaguar (Panthera onca palustris) with cub (estimated age 5 months), resting on a fallen tree over the Cuiaba River. Porto Jofre, Northern Pantanal, Mato Grosso State, Brazil, South America
Jaguar (Panthera onca) resting on river bank, Pantanal, Brazil
Water hyacinth (Eichhornia sp. ) and red Ludwigia (Ludwigia sp. ) in a clear stream, Pantanal, BrazilWater hyacinth (Eichhornia sp.) and red Ludwigia (Ludwigia sp.) in a clear stream, Pantanal, Brazil. Photographed for the Freshwater Project