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Tibetan antelope (Pantholops hodgsoni), two males, Kekexili, Qinghai, Tibetan Plateau, China, December
American bison (Bison bison) walking through snow using tracks made by other animals, Yellowstone National Park, Wyoming, USA, February
Bison (Bison bison) on frost covered ground, Yellowstone National Park, USA, February
Bison (Bison bison) close up head portrait with snow on fur, Yellowstone National Park, USA, February
Chamois (Rupicapra rupicapra), Ballons des Vosges Regional Natural Park, Vosges, Lorraine, France, December
Pyrenean chamois {Rupicapra rupicapra pyrenaica} Pyrenees, France
Cows (Bos taurus) in field of buttercups looking over dry stone wall. Peak District National Park, Derbyshire, UK, May 2012
American Bison (Bison bison) in habitat, Grand Teton National Park, Wyoming, USA, October
Female Ibex (Capra ibex) portrait, Creux du Van Natural Reserve, Val de Travers, Switzerland, December
Wildebeest (Connochaetes taurinus) herd walking in a line in the rain at sunset, Masai-Mara Game Reserve, Kenya. September
Topi (Damaliscus korrigum / Damaliscus lunatus) territorial male standing on mound looking across savanna. Masai-Mara game reserve, Kenya
Musk ox (Ovibos moschatus) two in habitat, Wrangel Island, Far Eastern Russia, October
Springbok (Antidorcas marsupialis) grazing in front of sand dunes, Namib Naukluft Park, Namibia, June 2006
Musk ox (Ovibos moschatus) portrait of two standing closely, Wrangel Island, Far Eastern Russia, September
Aerial view of two Gemsbok (Oryx gazella) in sand dunes, Namib Desert, Namibia
Chamois (Rupicapra rupicapra) walking with the moon behind, Mercantour National Park, France, October
Wildebeest (Connochaetes taurinus) crossing the Mara river, Masai-Mara Game Reserve, Kenya
American Buffalo / Bison (Bison bison) grazing in open plains. Yellowstone National Park, Wyoming, October
Highland cow on heather moorland, Curbar Edge, Peak District National Park, Derbyshire. August
Wild yak (Bos mutus) running, Kekexili, Qinghai, Tibetan plateau, China, December
Tibetan antelope (Pantholops hodgsonii) herd in the early morning with a low moon, Kekexili, Qinghai, Tibetan plateau, China, January
Alpine ibex (Capra ibex) adult male chasing two young males away, Gran Paradiso National Park, Italy, July
Domestic cows (Bos taurus) and free roaming horses grazing in the Pyrenees-Atlantiques, Pyrenees, France, June
Bison (Bison bison) in snowstorm. Yellowstone National Park, USA, February
Bison (Bison bison) two males head to head, Yellowstone NP, Wyoming, USA
Domestic sheep (Ovis aries) flock of Northumberland blackface sheep in snow, Tarset, Hexham, Northumberland, UK, November 2010
Blue sheep (Pseudois nayaur) Hemis National Park, Jammu and Kashmir, India
Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June
Saiga antelope (Saiga tatarica) male, standing in grassland in morning sunlight, Betpak Dala, Kazakhstan. Critically endangered
Domestic yak (Bos grunniens) lying under prayer flags near shores of Nam-Tso Lake (at 4, 730m), second largest salt lake in China. Tibet, China
Chamois (Rupicapra rupicapra) head portrait, Schwyz, Switzerland. November
Ibex (Capra ibex) feeding on grass, head portrait, Schwyz, Switzerland. August
Two Ibex (Capra ibex) males, standing on hillside, Lucerne, Switzerland. August
Iberian ibex (Capra pyrenaica), pair with male top right, during rutting season. Sierra Nevada National Park, Andalusia, Spain. October
Iberian ibex (Capra pyrenaica) kid peering over rocks. Sierra Nevada National Park, Andalusia, Spain. June
Iberian ibex (Capra pyrenaica) yearling making its way down snow covered slope. Sierra Nevada National Park, Andalusia, Spain. February
Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), female, being pestered by kid, aged two months, begging for milk. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica) kid, aged two months, making its way down cliff, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), female, grazing, being pestered by kid, aged two months, begging for milk, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July
Four Iberian ibex (Capra pyrenaica) kids, aged two months, playing, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), adult male, sitting down in snow covered landscape during snowfall. Sierra Nevada National Park, Andalusia, Spain. April
Iberian ibex (Capra pyrenaica), alert female with kid aged between one and two months. Sierra Nevada National Park, Andalusia, Spain. May
Iberian ibex (Capra pyrenaica), adult female, walking along coastal cliff. Mediterranean coast, Andalusia, Spain. November
Nilgiri tahr (Nilgiritragus hylocrius) standing next to road signage meant to warn vehicles about the possible road crossings of tahrs in the area, Valparai, Tamil Nadu, India
Springbok (Antidorcas marsupialis) herd in grassland after rain, Palmwag Conservancy, Damaraland, Namibia
Gemsbok (Oryx gazella) herd in desert after the wet season with Naukluft Mountains in the background, Namib-Naukluft National Park, Namibia
Gemsbok (Oryx gazella) pair standing in grass after wet season in desert, Namib-Naukluft National Park, Namibia
