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Impala antelope (Aepyceros melampus) close up of face, iMfolozi National Park, South Africa
Greater kudu (Tragelaphus strepsiceros) female running, St Lucia Wetlands National Park, South Africa
European bison / Wisent (Bison bonasus) herd released into the Tarcu mountains nature reserve, Natura 2000 area, Southern Carpathians, Romania. May 2014
RF- Arabian Oryx (Oryx leucoryx) Dubai Desert Conservation Reserve, Dubai, UAE
Iberian ibex (Capra pyrenaica) amongst scrub, Cliffs of Maro-Cerro Gordo Natural Area, Granada, Andalusia, Spain. January
Sichuan / Tibetan takin (Budorcas taxicolor tibetana) young calf that lost his mother, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
African lioness (Panthera leo) sitting patiently in the long grass, watching herd of Common eland (Tragelaphus oryx). Focus on lion. Grumeti Reserve, Northern Tanzania
Musk ox (Ovibos moschatus) two in profile, Wrangel Island, Far Eastern Russia, September
Musk ox (Ovibos moschatus) at sunset, Wrangel Island, Far Eastern Russia
Blackbuck (Antelope cervicapra), female, running with high jumps known as Pronking. Tal Chhapar Wildlife Sanctuary, Rajasthan, India
Wild yak, (Bos mutus), Keke Xili, Changtang, Tibetan Plateau, Qinghai, China
Wild yak herd (Bos mutus) Keke Xili, Changtang, Tibetan Plateau, Qinghai, China
BlackbuckA(Antilope cervicapra), herd with males and females, Velavadar National ParkBlackbuck(Antilope cervicapra), herd with males and females, Velavadar National Park, Gujarat, India
Nilgai (Boselaphus tragocamelus), female in water, Rajasthan, India
Bonnet macaque (Macaca radiata), young animal on fence, Karnataka, India
Greater blue sheep (Pseudois nayaur), herd on mountain slope, important prey for snow leopards (Panthera uncia), Himalaya, Hemis National Park, Ladakh, India
RF -Three Bison (Bison bison) walking through snow with herd feeding in background. Yellowstone National Park, Wyoming, USA. January
Bison (Bison bison) standing in geothermal run-off in winter, Yellowstone National Park, Wyoming, USA, February
Bighorn Sheep (Ovis canadensis) on the hillside, ruminating, Yellowstone National Park, Wyoming, USA. January 2011
Bison (Bison bison) standing steam from geothermal springs, Yellowstone National Park, Wyoming, USA, February
RF- Bison (Bison bison) standing in steam from geothermal springs. Yellowstone National Park, Wyoming, USA, February
Male impalas (Aepyceros melampus) standing in heavy rain, Masai-Mara Game Reserve, Kenya. February
Portrait of male Waterbuck (Kobus ellipsiprymnus) in the rain, Masai-Mara game reserve, Kenya
Wildebeest (Connochaetes taurinus) herd below stormy sky during migration, Masai-Mara game reserve, Kenya
Lioness (Panthera leo) and Thomsons gazelles (Eudorcas thomsonii) silhouetted at sunset, Masai-Mara game reserve, Kenya
Red lechwe (Kobus leche) silhouetted at sunrise, Okavango Delta, Botswana
Musk ox (Ovibos moschatus) portrait, Wrangel Island, Far Eastern Russia, March
European bison (Bison bonasus), Drawsko Military area, Western Pomerania, Poland, February
Herd of Impala (Aepyceros melampus) silhouetted at sunset, Ngorongoro Conservation Area, Tanzania
Snow Leopard (Uncia uncia) with Himalayan Blue Sheep (Pseudois nayaur) kill, Hemas National Park, Ladakh, India
Two male Alpine ibex (Capra ibex ibex) fighting in front of the Mer de Glace glacier, with a female watching, Aiguilles Rouges (Red Peaks) Regional Natural Park, Haute-Savoie, France, June
Impala (Aepyceros melampus) females, Masai-Mara Game Reserve, Kenya
Mature Bighorn Rams (Ovis canadensis) on high mountain pass. Jasper National Park, Alberta, Canada, October
Blue Sheep (Pseudois nayaur) on hillside, Kekexili, Qinghai, China, January
American Bison (Bison bison) walking through snow field, Hayden Valley, Yellowstone National Park, Wyoming, USA, January
American Bison (Bison bison) lying in snow field, Hayden Valley, Yellowstone National Park, Wyoming, USA, January
Springbok (Antidorcas marsupialis) herd, Kgalagadi Transfrontier Park, South Africa
Bighorn sheep (Ovis canadensis) male ram walking in the snow, Yellowstone National Park, USA, February
Bison (Bison bison) herd walking in line in snow, Yellowstone National Park, Wyoming, USA, February
Bison (Bison bison) single animal in snow, Yellowstone National Park, Wyoming, USA, February
Alpine ibex (Capra ibex ibex) in front of the Mont Blanc massif, seen from the Aiguilles Rouges (Red Peaks) Regional Natural Park, Haute-Savoie, France, June
Male Ibex (Capra ibex), Aiguilles Rouges (Red Peaks) National Nature Reserve, Haute-Savoie, France, June
Grants zebra (Equus burchelli boehmi) in a migration herd of wildebeest (Connochaetes taurinus) Masai-Mara Game Reserve, Kenya
Gemsbok / South African oryx (Oryx gazella) on sand dune, Namib Desert, Sossusvlei, Namibia, October 2015
Buffons kob (Kobus Kob) on track in Pendjari National Park, Benin
Urial sheep (Ovis vignei) herd running across steep barren slopes. Himalayas near Ulley, Ladakh, India
Thomsons gazelle (Eudorcas thomsonii) herd with Wildebeest herd visible beyond, Masai-Mara game reserve, Kenya