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Mist hanging over Lowland Dipterocarp Rainforest just after sunrise. Danum Valley Conservation Area, Sabah, Borneo
Two Millipedes (Julidae) close up portrait, Mulu, Sarawak, Malaysia
Great anglehead lizard (Gonocephalus grandis) resting on tree trunk, Mulu, Sarawak, Malaysia
Black-headed cat snake (Boiga nigriceps) coiled, portrait, Kubah National Park, Sarawak, Malaysia
Male Gold-specs jawfish (Opistognathus randalli), with mouth brooding eggs, the eyes can clearly be seen in the eggs indicating they will soon hatch, Mabul Island, Sabah, Malaysia, Celebes Sea
Swarm of Asian weaver ants (Oecophylla smaragdina) attacking a Finger-print ant (Diacamma sp)Swarm of Asian weaver ants (Oecophylla smaragdina) attacking a Finger-print ant (Diacamma sp.), Kubah National Park, Sarawak, Borneo
Close up of hands of a Bornean Orangutan (Pongo pygmaeus), Tanjung Puting National Park, Borneo
Orange backed red langur (Presbytis rubicunda chrysea) in rainforest understorey. Danum Valley, Sabah, Borneo, Malaysia
Carrion fly on the parasitic plant Rafflesia pricei flower, Mount Kinabalu, Borneo, May, 2013. Filmed for the BBC series Wonders of The Monsoon
Montane horned frog (Megophrys kobayashii) head close up, Kinabalu National Park, Sabah, Borneo, Malaysia
Minute / Dwarf filefish (Rudarius minutus) just over an inch long as an adult, Celebes Sea, Mabul Island, Malaysia
Sunda leopard cat (Prionailurus javanensis) kitten. Sabah, Malaysian Borneo
Blyths paradise flycatcher (Terpsiphone affinis) male calling, Sabah, Malaysian Borneo
Proboscis monkey (Nasalis larvatus) male leaps between trees. Sabah, Malaysian Borneo
Lowland dipterocarp rainforest, with huge Tualang (Tapang) or Mengaris Tree (Koompassia excelsa) in the foreground, Danum Valley, Sabah, Borneo
Western / Horsfields tarsier (Tarsius bancanus) hunting invertebrate prey in rainforest understory vegetation at night. Danum Valley, Sabah, Borneo
Giant shield bug nymphs (Pycanum sp. ), recently hatched from eggsGiant shield bug nymphs (Pycanum sp.), recently hatched from eggs. Danum Valley, Sabah, Borneo
Safety in Numbers - Giant Shield bug nymphs (Pycanum sp.), recently hatched from eggs, with a final individual still to emerge
Arboreal tarantula (Lampropelma sp. ) female guarding spiderlings at entrance to hollowArboreal tarantula (Lampropelma sp.) female guarding spiderlings at entrance to hollow retreat, 4 metres up a tree. Danum Valley, Sabah, Borneo
Lantern bug (Pyrops whiteheadi), detail of wing. Danum Valley, Sabah, Borneo
Lantern bug (Pyrops whiteheadi), detail of head. Danum Valley, Sabah, Borneo
Lowland dipterocarp rainforest at dawn, Danum Valley, Sabah, Borneo
Sumatran pitviper (Trimeresurus sumatranus), detail of scales. Danum Valley, Sabah, Borneo
RF - Maliau Falls, Maliau River. Centre of Maliau Basin - Sabah, Borneo. (This image may be licensed either as rights managed or royalty free.)
Portrait of Bornean Horned Frog (Megophrys nasuta) amongst leaf-litter in forest floor. Taken at night. Danum Valley, Sabah, Borneo
Crinoid (Crinoidea) on Soft coral (Alcyonacea). Derawan Islands, East Kalimantan, Indonesia
Pink anemonefish (Amphiprion perideraion). Derawan Islands, East Kalimantan, Indonesia
Close up of decaying leaf from rainforest floor, showing lichens and moulds. Danum Valley, Sabah, Borneo
One of the dams constructed by the local field staff of the Borneo Nature Foundation in the canals in the Sabangau (peat-swamp) Forest. These are made to combat forest fires
Pandan (Pandanus sp. ) growing in the Sabangau River, Central Kalimantan, IndonesiaPandan (Pandanus sp.) growing in the Sabangau River, Central Kalimantan, Indonesia
Maroon / Red leaf monkey / Langur (Presbytis rubicunda) eating fruits in tree, Danum Valley Conservation Area, Sabah, Borneo, Malaysia
Red leaf monkey (Presbytis rubicunda) female feeding in Strangler fig tree (Ficus dubia) Gunung Palung National Park, Kalimantan, Borneo, Indonesia
Crocodilefish (Cymbacephalus beauforti) close up detail of the retractable eye-lashes, Mabul Island, Sabah, Borneo, Malaysia Celebes Sea
RF - Camouflaged moth (Agathia codina) Kinabalu National Park, Sabah, Malaysian Borneo. September
Moth (Eumelea sp) Danum Valley, Sabah, Borneo
Bornean orangutan (Pongo pygmaeus) mother and baby, Tanjung Puting National Park, Borneo-Kalimatan, Indonesia, endangered species
Orang utan (Pongo pygmaeus) head portrait of female, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Sarawak, Borneo, Malaysia, Endangered
Orang utan (Pongo pygmaeus) portrait, swinging, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
Fungus gnat (Mycetophilidae) larvae, with sticky hanging threads, waiting to ambush small flying insects. Deer caves, Gunung Mulu National Park, Borneo, Sarawak, Malaysia
Komodo Dragon (Varanus komodoensis) close up, Komodo National Park, Indonesia
Rhinoceros beetle (Chalcosoma moellenkampi) climbing up tree trunk, Danum Valley Reserve, Sabah, Borneo, Malaysia
RF- Sunrise and mist over lowland dipterocarp rainforest. Danum valley, Sabah, Borneo, Malaysia
Spiky flower-mimic stick insect (Toxodera berieri) active at night. Danum Valley, Sabah, Borneo
Lowland dipterocarp rainforest at dawn. Danum Valley, Sabah, Borneo, May 2011
Long-tailed / Crab-eating macaque (Macaca fascicularis) foraging on coastline at low tide, Bako National Park, Sarawak, Borneo, Malaysia
Proboscis Monkey (Nasalis larvatus) female sitting in tree, partially obscured by soft focus leaves in foreground, Bako National Park, Sarawak, Borneo, Malaysia
Eye and spiracle detail of a Bluespotted / Ribbontail Stingray (Taeniura lymma) Sipadan Island, Sabah, Borneo, Malaysia. May
Lowland dipterocarp rainforest canopy at dawn. Danum Valley, Sabah, Borneo, May 2011