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Large shoal of Horse mackerel (Megalaspis cordyla) schooling. Rinca, Komodo National Park, Indonesia
Magnificent sea anemone (Heteractis magnifica) with Anemone fish within tentacles, Indonesia. Indo-Pacific
Blue butterfly (Lycaenidae) frozen in ice, Cortes de la Frontera, Los Alcornocales Natural Park. Spain
Victoria crowned pigeon (Goura victoria) head portrait, vulnerable species, captive
Thread-tail dartgoby (Ptereleotris hanae) Sharp Island or Kiu Tsui Chau, Kiu Tsui Country Park, Hong Kong, China
Ocean sunfish (Mola mola) off Halifax, Nova Scotia, Canada. July
Atlantic wolfish (Anarhichas lupus) looking out hole, Bay of Fundy, New Brunswick, Canada, July
Blue poppy (Meconopsis horridula) Sanjiangyuan National Nature Reserve, Qinghai Hoh Xil UNESCO World Heritage Site, Qinghai-Tibet Plateau, Qinghai Province, China
Bioluminescent Sea-fireflies (Vargula hilgendorfii) producing a bright blue light. The light is produced by mixing two chemicals together in the presence of oxygen
Portrait of a Golden snub-nosed monkey 1+Rhinopithecus roxellana+2 full frame of the face, Foping Nature Reserve, Shaanxi, China. Endangered species
Blue shark (Prionace glauca) off Halifax, Nova Scotia, Canada. July
Blue whale (Balaenoptera musculus) diving beneath the surface. Indian Ocean, Sri Lanka
Brydes whale (Balaenoptera edeni) Trincomalee, Eastern Province, Sri Lanka, Bay of Bengal, Indian Ocean
RF - Green-and-Black Poison Dart Frog (Dendrobates auratus) inside bromiliad. Boca Tapada, Caribbean slope, Costa Rica, Central America
Bioluminescent Fungus gnat (Arachnocampa luminosa) larva attached to the cave roof with silk threads, Glowworm cave near Waitomo Cave, near Te Kuiti, North Island, New Zealand, July
Quinling Golden snub nosed Monkey (Rhinopitecus roxellana qinlingensis), adult male face portrait. Zhouzhi Nature Reserve, Qinling Mountains, Shaanxi, China
Pine tree at night with moon shining, on Stuoc peak, Durmitor NP, Montenegro, October 2008
Dead Pine trees at night on Stuoc peak, Durmitor NP, Montenegro, October 2008
Atlantic bluefin tuna (Thunnus thynnus) portrait, captive, Malta, Mediteranean, May 2009 WWE BOOK
Arctic tern (Sterna paradisaea) in flight at twilight using slow shutter speed to accentuate blur and motion in low-key light, Inner Farne, Farne Islands, Northumberland, July
Oystercatcher (Haematopus ostralegus) at dusk, with lights in distance, South Swale, Kent, England, UK, December. 2020VISION Book Plate
RF- Rear view of Basking shark (Cetorhinus maximus) feeding on plankton, visible as white dots on water surface near island of Coll, Inner Hebrides, Scotland, UK. June
Golden Retriever puppy in blue flowers. USA
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea in the late afternoon, close to Little Brother Island
Waves lashing the Four Lighthouse during winter storm, Northern Brittany, France, December 2011. Sequence 8/12
Orcas / Killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Lilac-breasted roller (Coracias caudata) stretching wings on acacia branch, Masai Mara Game Reserve, Kenya. November
Salt marsh cord grass (Spartina alterniflora) on shore of Cape Cod, with Long Point Lighthouse in the background, Massachusetts, USA, September
Geranium Argus butterfly (Plebejus eumedon) feeding on Wood Cranesbill (Geranium sylvaticum) northern Finland, June
Lesser flamingo (Phoeniconaias minor) portrait, Lake Bogoria, Rift valley, Kenya, Africa
Fringed leaf frog (Cruziohyla craspedopus) on branch, Yasuni National Park, Orellana, Ecuador
Rangimairewhenua or Blue Lake, Nelson Lakes National Park, Southern Alps, New Zealand, February 2013. Photographed for The Freshwater Project
RF -Three colourful Red-and-green macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
RF - Four colourful Red-and-green macaws or Green-winged macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
Red-and-green macaw (Ara chloropterus) in flight over forest canopy. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
European / Common Lobster (Homarus gammarus) in rocky retreat. Controlled conditions, Cornwall, UK. May
Silver-studded blue butterflies (Plebejus argus) resting on grass and backlit by the sun, Upton Towans, Cornwall, UK. June 2017
Kingfisher (Alcedo atthis) male passing fish to female spring courtship behaviour, Hertfordshire, England, UK, March. Sequence 4 of 6
School of large Atlantic bluefin tuna (Thunnus thynnus) captive in growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Hooded nudibranchs (Melibe leonina) filter feeding beneath bull kelp. Hussar Point, Browning Pass, Port Hardy, Vancouver Island, British Columbia, Canada
RF - Close up detail of the eye and skin pattern of a Giant hawkfish (Cirrhitus rivulatus). Socorro Island, Pacific Ocean
Male Great white shark (Carcharodon carcharias) with sunrays, Guadalupe Island, Mexico, Pacific Ocean
Meadow Cranesbill (Geranium sanguineum) in flower. Germany, May
Black eyed susan (Rudbeckia fulgida var. sullivantii) Goldsturm flowers with Michaelmas daisies (Aster amellus) in garden
Palm tree branch dipping towards water, Puerto Jimenez, Golfo Dulce, Osa Peninsula, Costa Rica
Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
Brick wall in the Blue City, Jodhpur, Rajasthan, India
Blue and gold fusilier / Scissor tailed fusilier (Caesio caerulaurea) school swimming through shafts of sunlight, West Papua, Indonesia
