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Sparkling violetear hummingbird (Colibri coruscans) incubating eggs at nest, Andean cloud forest, Ecuador
Plate billed mountain toucan (Andigena laminirostris) Bellavista cloud forest private reserve, 1700m altitude, Tandayapa Valley, Andean cloud forest, Ecuador
Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019
Corncrake (Crex crex) walking at dawn on Machair, North Uist, Scotland, UK, June
Two Little terns (Sterna albifrons) at the nest amongst Black oats (Avena strigosa) growing on shell rich sands Machair, North Uist, Scotland, UK, June
Little tern (Sterna albifrons) at the nest amongst Black oats (Avena strigosa) growing on shell rich sands Machair, North Uist, Scotland, UK, June
Little tern (Sterna albifrons) with newly hatched chick, Gronant dunes, Denbighshire, Wales, UK, June
Little tern (Sterna albifrons) nest with three eggs in shingle ridge. County Wicklow, Ireland, June
James flamingo / Puna flamingo (Phoenicoparrus jamesi). Lago Colorado, Bolivia. March
King penguins (Aptenodytes patagonicus) walking in line on a windy beach. Sanders Island, Falklands. November
Snowy egret (Egretta thula) shows its breeding plumage. St. Johns Management Area, Florida, USA, March
Yellow warbler (Setophaga petechia aureola) on lava rocks. Isabela Island, Galapagos. May
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Western screech owl (Megascops kennicottii) is camouflaged as it looks from a tree cavity. Bosque del Apache National Wildlife Refuge, New Mexico, USA, January
Blue tit (Cyanistes caeruleus) bringing Crab spider (Misumena vatia) prey to nest, Monmouthshire Wales, UK, May
Pectoral sandpiper (Calidris melanotos) juvenile feeding around edge of shallow flood, Llanrhystud, Ceredigion, West Wales, UK, September
Rock pipit (Anthus petrosus) foraging for invertebrates among seaweed on the strand line as the tide rises at sunset, Cornwall, UK, September
Flock of Oystercatchers (Haematopus ostralegus) and Knot (Calidris canuta) in flight and on tidal shallows. The Wash Estuary, Norfolk, September
Barn Swallow (Hirundo rustica) feeding a fledgling on a wire. Perthshire, Scotland, September
Sparrowhawk (Accipiter nisus) watching for prey from a garden rowan tree. Perthshire, Scotland, September
Oystercatcher (Haematopus ostralegus) flock in flight, Liverpool Bay, Lancashire, UK, September
Group of Greylag geese (Anser anser) flying in the half light before dawn, Snettisham RSPB reserve, Norfolk, UK, September
RF - Black Grouse (Tetrao tetrix) male peforming flutter jump display on lek, Cairngorms National Park, Scotland, UK.May (This image may be licensed either as rights managed or royalty free.)
Crested tit (Parus cristatus) backlit on lichen-covered branch, Inshriach Forest, Scotland, UK, January.January
Turnstones (Arenaria interpres) and Purple sandpipers (Calidris maritima) over wintering on the beaches of the Orkney Isles, Scotland, UK, April
Aerial view of nesting Gannets (Morus bassanus) and swirling seas, Hermaness, Shetland, Scotland, UK, October
Great bittern (Botaurus stellaris) head and neck, lookin up through reeds. Suffolk, UK. November
Great bittern (Botaurus stellaris) portrait. Suffolk, UK. November
Darter ( Anhinga melanogaster) drying wings, Keoladeo NP, Bharatpur, India
Olive-backed sunbird (Nectarinia jugularis) female returning to her nest with nesting material, Cape Hillsborough National Park, Queensland, Australia, September
Emu (Dromaius novaehollandiae) juvenile with beak open to catch a leaf, Cleland Wildlife Park, South Australia, March. Captive
Gannet (Morus bassanus) diving for fish, Noss, Shetland Islands, Scotland, UK, August
Egyptian goose (Alopochen aegyptiacus) in flight over Rutland Water, Rutland, UK, August
Swallows (Hirundo rustica) group of four chicks waiting to be fed, Norfolk, England, UK, August
Common tern (Sterna hirundo) in flight, Norfolk, UK, August
Gannet (Morus bassanus) adult, backlit by the morning sun, in flight. Shetland Islands, Scotland, UK. August
Grey heron (Ardea cinerea) head profile, Regents Park, London, UK August
Kittiwake (Rissa tridactyla) in flight, juvenile. Bempton, Yorkshire, UK, August
Red-necked phalarope (Phalaropus lobatus) female about to strike at a fly hovering nearby. Shetland Islands, Scotland, UK, August (non-ex)
Golden plover (Pluvialis apricaria) profile portrait on open moorland in evening light. Shetland Islands, Scotland, UK, August (non-ex)
Andean Emerald Hummingbird (Amazilia franciae), Mindo cloud forest area, Ecuador, July
Mandarin duck drake (Aix galericulata) floating on colourful water, Southwest London, UK, January
Sword-billed hummingbird (Ensifera ensifera) in flight, North-Ecuador, Ecuador
Grey-headed woodpecker (Picus canus) on tree trunk in falling snow. Kalvtrask, Vasterbotten, Lapland, Sweden. January
Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July
Shoebill stork (Balaeniceps rex) in the swamps of Mabamba, Lake Victoria, Uganda, February
Dunlin (Calidris alpina) portrait. Vardo, Finnmark, Norway. May
Black woodpecker (Dryocopus martius) male perched on tree trunk, trees covered in hoar frost. Kuusamo, Northern Ostrobothnia, Finland. February